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Verbena litoralis

Kunth (1818)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Lamiales Family: Verbenaceae Genus: Verbena

‘ōwī [owi], ha‘uoi (Ni‘ihau via St. John), oī [oi], ōwī, vervain

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Description

Key Characters:

Growth Form: Erect or ascending perennial herbs 4–20 dm tall, sometimes somewhat woody toward base.

Stems: Stems few to many-branched, glabrous or sparsely strigillose.

Roots:

Leaves: Leaves simple. Opposite and decussate (pairs of leaves alternating 90 degrees between nodes). Blades lanceolate to oblanceolate, 3–10 cm long, 1–1.5 cm wide. Apex acute. Base attenuate. Surfaces finely scaberulous. Margins coarsely serrate. Veins prominent. Sessile. Stipules absent.

Flowers: Flowers numerous in several slender, terminal spikes 3–12 cm long, sparsely short–pilose throughout; Bracts subulate, shorter than to as long as the calyx; short–pilose. Flowers bisexual (perfect), alternate, nearly zygomorphic. Calyx sometimes purple, 1.5–2.3 mm long. Corolla usually blue, occasionally pale blue, lavender, or purple, ca. 2–3 mm long, 5-lobed, imbricate. Stamens 4, in 2 pairs, inserted on and included in corolla tube; anthers dithecal, opening by longitudinal slits. Ovary superior, 4-lobed, 2-carpellate, 4-celled, placentation axile; ovules 1 per cell, attached laterally or near base, erect, anatropous; style 2-lobed, the lobes dissimilar, only 1 stigmatic.

Fruit: Schizocarps readily separating at maturity into 4 crustaceous nutlets; pericarp hard and dry. Seeds 1 per nutlet; oblong; triangular in cross section; endosperm essentially absent.

Ploidy: 2n = 28; 56

Habitat: Occurring in dry to wet; disturbed habitats.

Elevation Range: 10–2,280 m.

Historical Distribution

Images

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Synonyms (32)

Verbena affinis M.Martens & Galeotti (1844)
Verbena bonariensis (Misapplied)
Verbena bonariensis var. litoralis (Kunth) Hook. ex Müll.Berol. (1845)
Verbena brasiliensis var. subglabrata Moldenke (1950)
Verbena cajamarcensis Binder ex Molinari (2015)
Verbena caracasana Kunth (1818)
Verbena carolina var. glabra Moldenke (1959)
Verbena gentryi Moldenke (1941)
Verbena glabrata var. tenuispicata Moldenke (1967)
Verbena integrifolia Sessé & Moc. (1886)
Verbena integrifolia f. albiflora Moldenke (1968)
Verbena lanceolata Willd. ex Spreng. (1825)
Verbena litoralis f. albiflora (Moldenke) Moldenke (1979)
Verbena litoralis f. magnifolia Moldenke (1977)
Verbena litoralis var. albiflora Moldenke (1940)
Verbena litoralis var. caracasana (Kunth) Briq. (1904)
Verbena litoralis var. glabrior Benth. (1846)
Verbena litoralis var. leptostachya Schauer (1847)
Verbena litoralis var. litoralis
Verbena litoralis var. melanopotamica Hauman (1913)
Verbena litoralis var. portoricensis Moldenke (1982)
Verbena litoralis var. pycnostachya Schauer (1847)
Verbena litoralis var. subglabrata (Moldenke) N.O'Leary & Múlgura (2012)
Verbena longifolia M.Martens & Galeotti (1844)
Verbena longifolia f. albiflora Moldenke (1961)
Verbena longifolia var. pubescens Moldenke (1966)
Verbena minutiflora var. peruviana Moldenke (1981)
Verbena nudiflora Nutt. ex Turcz. (1863)
Verbena paucifolia Turcz. (1863)
Verbena sedula Moldenke (1955)
Verbena sedula var. darwinii Moldenke (1968)
Verbena sedula var. fournieri Moldenke (1969)

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Ni'ihau(Incl. Lehua) Naturalized
Kuaihelani (Midway Atoll) Naturalized
Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Lana'i Naturalized
Kaho'olawe Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Notes

  • Description digitized by Ikaika Mendez
  • Description digitized from the Manual of the Flowering Plants of Hawaii
  • Has been used medicinally in Hawai'i especially as a mash applied to cuts and bruises and also to sprained and fractured areas.
  • in 1837, specific locality unknown

Bibliography

Name Published In: F.W.H.von Humboldt, A.J.A.Bonpland & C.S.Kunth, Nov. Gen. Sp. 2: 276 (1818)

Other References

Wagner et al. 1990:1325 (Mi, Ni, K, O, Mo, L, M, Ka, H)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
2 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
3 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
4 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
5 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
6 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
7 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
8 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
9 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
10 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
11 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
12 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
13 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
14 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
15 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
16 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
17 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
18 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
19 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
20 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
21 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
22 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
23 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
24 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
25 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
26 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
27 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
28 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
29 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
30 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
31 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
32 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
33 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
34 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
35 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
36 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
37 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
38 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
39 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
40 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
41 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
42 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
43 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
44 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
45 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
46 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
47 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
48 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
49 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
50 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
51 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
52 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
53 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
54 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
55 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
56 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
57 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
58 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
59 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
60 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
61 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
62 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
63 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
64 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
65 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
66 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
67 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
68 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
69 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
70 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
71 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
72 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
73 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
74 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
75 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
76 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
77 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
78 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
79 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
80 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
81 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
82 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
83 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
84 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
85 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
86 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
87 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
88 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
89 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
90 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
91 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
92 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
93 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
94 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
95 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
96 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
97 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
98 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
99 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
100 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
101 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
102 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
103 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
104 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
105 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
106 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
107 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
108 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
109 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
110 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
111 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
112 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
113 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
114 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
115 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
116 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
117 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
118 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
119 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
120 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
121 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
122 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
123 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
124 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
125 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
126 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
127 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
128 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
129 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
130 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
131 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
132 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
133 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
134 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
135 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
136 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
137 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
138 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
139 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
140 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
141 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
142 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
143 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
144 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
145 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
146 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
147 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
148 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
149 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
150 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
151 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
152 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
153 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
154 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
155 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
156 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
157 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
158 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
159 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
160 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
161 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
162 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
163 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
164 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
165 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
166 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
167 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
168 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
169 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
170 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
171 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
172 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
173 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
174 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
175 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
176 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
177 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
178 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
179 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
180 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
181 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
182 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
183 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
184 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
185 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
186 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
187 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
188 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
189 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
190 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
191 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
192 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
193 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
194 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
195 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
196 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
197 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
198 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
199 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
200 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
201 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
202 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
203 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
204 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
205 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
206 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
207 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
208 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
209 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
210 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
211 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
212 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
213 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
214 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
215 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
216 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
217 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
218 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
219 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
220 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
221 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
222 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
223 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
224 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
225 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
226 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
227 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
228 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
229 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
230 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
231 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
232 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
233 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
234 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
235 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
236 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
237 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
238 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
239 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
240 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
241 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
242 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
243 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
244 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
245 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
246 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
247 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
248 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
249 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
250 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
251 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
252 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
253 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
254 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
255 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
256 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
257 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
258 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
259 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
260 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
261 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
262 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
263 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
264 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
265 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
266 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
267 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
268 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
269 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
270 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
271 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
272 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
273 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
274 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
275 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
276 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
277 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
278 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
279 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
280 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
281 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
282 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
283 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
284 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
285 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
286 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
287 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
288 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
289 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
290 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
291 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
292 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
293 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
294 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
295 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
296 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
297 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
298 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
299 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
300 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
301 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
302 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
303 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
304 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
305 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
306 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
307 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
308 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
309 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
310 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
311 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
312 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
313 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
314 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
315 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
316 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
317 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
318 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
319 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
320 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
321 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
322 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
323 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
324 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
325 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
326 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
327 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
328 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
329 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
330 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
331 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
332 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
333 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
334 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
335 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
336 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
337 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
338 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
339 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
340 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
341 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
342 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
343 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
344 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
345 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
346 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
347 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
348 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
349 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
350 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
351 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
352 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
353 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
354 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
355 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
356 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
357 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
358 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
359 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
360 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
361 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
362 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
363 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
364 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
365 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
366 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
367 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
368 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
369 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
370 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
371 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
372 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
373 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
374 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
375 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
376 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
377 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
378 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
379 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
380 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
381 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
382 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
383 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
384 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
385 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
386 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
387 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
388 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
389 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
390 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
391 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
392 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
393 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
394 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
395 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
396 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
397 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
398 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
399 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
400 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
401 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
402 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
403 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
404 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
405 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
406 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
407 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
408 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
409 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
410 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
411 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
412 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
413 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
414 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
415 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
416 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
417 Verbena litoralis Koloa, Kalaheo, upper end of Puuwai Rd. wet roadside secondary forest; Wedelia Preserved_Specimen to 0.5 m herb, slender; square stem, small purple flowers Trauernicht, C. 205 Kauai BISH 2007-10-15
418 Verbena litoralis Nawaimaka drainage. Acacia-Metrosideros mesic forest, open to closed canopy, riparian with moderate to steep banks. W/Kadua affinis, Dodonaea, Leptecophylla tameiameiae, Psychotria mariniana, Antidesma platyphylla, Pouteria, Xylosma hawaiiense, Alyxia stellata, Chamaesyce atrocicca, Carex alligata, Cyrtandra kauai Preserved_Specimen 1.5 m. tall, stems green, corolla pale pruple. Wood, K.R. 13816 Kauai BISH 2009-07-16
419 Verbena litoralis Kau District: Keauhou Ranch Along fenceline of horse corrall in lower pastures of ranch. Preserved_Specimen Herb, 3 ft. tall, purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
420 Verbena litoralis stream bank below Dillingham Field wetland grasses Preserved_Specimen Funk, E.J. 1105 Oahu BISH 1990-05-08
421 Verbena litoralis ridge north of Kuapa Pond open, hillsides Preserved_Specimen Gillespie, J.W. 1076 Oahu BISH 1927-07-08
422 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
423 Verbena litoralis Makanikahau Reservoir, Old Dried mud Preserved_Specimen St.John, H. 23633 Niihau BISH 1949-04-01
424 Verbena litoralis Puu Huluhulu. Near the escarpment Transect (T16) Remnant dryland community. Common subshrub among Pennisetum setaceum. Preserved_Specimen Common subshrub. Takeuchi, W.N. 5714 Hawaii BISH 1989-12-26
425 Verbena litoralis Keauhou Ranch, Kau District Alogn fenceline of horse corrall. Preserved_Specimen Herb, 3 ft. tall, with purple flowers. Davis, J. 166 Hawaii BISH 1979-12-18
426 Verbena litoralis Peacock Flat, Waianae Mts roadside, dry forest Preserved_Specimen herb to 1 m. Stems are quadrangular. Leaves opposite; elliptic-oblong; margins serrate; tops glandular, hirsute below. Flowering heads terminal, small. Flowers blue. Spence, G. 209 Oahu BISH 1973-09-06
427 Verbena litoralis Hawaii Volcanoes National Park. Along Kipuka Puaulu trail, to the right of no. 17 signpost in open sunny area. Preserved_Specimen Herat, T. 559 Hawaii BISH 1974-06-12
428 Verbena litoralis North Kona District; Burn area, ca 1.5 km E of Western end of Kona Highway, ca 300 m SW of Keamuku Flow, ca 300 m SSE of Boundary Pennisetum grassland, gentle slopes, A'a lava; associates Pennisetum setaceum, Dodonaea viscosa Preserved_Specimen Close, B. 109426 Hawaii BISH 1994-10-29
429 Verbena litoralis Kaluanui Flats Pineapple fields Preserved_Specimen Herb to 1 m; stem quadrangular. Leaves opposite, elliptic-oblong, margins serrate. Flowering terminal small with blue corollas. Spence, G. 369 Lanai BISH 1973-10-24
430 Verbena litoralis Keanakolu, Hilo Forest Reserve Vegetation: Dryopteris, Pennisetum clandestinum, Acacia koa, Coprosma, Passiflora mollissima and Rubus. Preserved_Specimen Higashino, P.K. 6965 Hawaii BISH 1978-03-01
431 Verbena litoralis Mauna Kea (Puu Laau Road) In open Mamane-Naio forest. Preserved_Specimen Engilis Jr., A. 39 Hawaii BISH 1987-08-19
432 Verbena litoralis Puu Laau Weed growing near the hunter's cabin. Preserved_Specimen Herbst, D.R. 5210 Hawaii BISH 1975-01-18
433 Verbena litoralis Laie clay, pasture Preserved_Specimen herb 1-6', tall slender stems Guinto, J. 27 Oahu BISH 1986-03-06
434 Verbena litoralis Kilauea weed in abandoned sorghum field Preserved_Specimen Herbst, D.R. 5244 Kauai BISH 1975-02-27
435 Verbena litoralis Kahaluu; Kailua; Kapua Koa forest. Preserved_Specimen Adee, K.T. Hawaii BISH
436 Verbena litoralis Puuwaawaa; track to 1986 burn area (from Ranch HQ) 4WD access track to burn area. Preserved_Specimen Common roadside weed. Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
437 Verbena litoralis Kawa side of Kahana Valley, pass the main gate and the marshy area along jeep trail, open areas Preserved_Specimen common weed, ~4', with light blue flowers Herat, T. 158 Oahu BISH 1972-08-14
438 Verbena litoralis Kahuku, Ocean View Estates, open dry forest on a'a lava at junction of King Kamehameha Blvd. and Walaka Drive. Preserved_Specimen Common roadside weed with blue flowers. Herbst, D.R. 5068 Hawaii BISH 1974-10-09
439 Verbena litoralis Kewalo, back of Punchbowl open, dry, roadside Preserved_Specimen 1 m tall; flower - small, inconspicous, lavender Judd, C.S. 11 Oahu BISH 1924-11-06
440 Verbena litoralis Honolulu, 3830 Tantalus Dr., access road to Puu Ohia roadside in wet forest; with Phyllostachys nigra, Bidens cynapiifolia, Panicum maximum. Preserved_Specimen to 4-20 dm tall; corolla blue Ray, G. 50 Oahu BISH 1997-09-15
441 Verbena litoralis Waikane-Schofield trail, Koolau Mts, Kahana along trail Preserved_Specimen fls. lavendar Fosberg, F.R. 10276 Oahu BISH 1933-09-22
442 Verbena litoralis 1750' Field 123, Honokaa GSugar Co. Hamakua District Common weed at field edge, and on jeep roads through field, not present inside mature stand. Preserved_Specimen Perennial root, plants 30- 48 tall, corolla pale blue Tomich, P.Q. 911 Hawaii BISH 1960-03-18
443 Verbena litoralis Lower half of Kaala Road mesic forest Preserved_Specimen flowers lavender Annable, C.R. 3727 Oahu BISH 1997-10-15
444 Verbena litoralis E central part of island, slope of Moaula Herb about 3' tall growing in shade of Prosopis pallida tree on rock outcropping. Preserved_Specimen Flowers purple. Cuddihy, L.W. 428 Kahoolawe BISH 1980-04-25
445 Verbena litoralis Trail to Kaluiwaa (Sacred Falls), outside forest Reserve At base of foot path trail. Preserved_Specimen Corn, C. Oahu BISH 1978-03-21
446 Verbena litoralis Drive down Chain of Craters Road about 12.9 miles toward Kalapana from intersection with Crater Rim Road Plant was across road (N) from turnoff to Naulu Picnic Area. (plot 5) Shrub about 3 feet tall, at roadside.. Preserved_Specimen Newell, C.L. 471 Hawaii BISH 1966-08-20
447 Verbena litoralis Pohakuloa on Mauna Kea; above Pohakuloa Military Camp. Pohakuloa burn 1979 Pohakuloa burn 1979. Preserved_Specimen Adee, K.T. Hawaii BISH
448 Verbena litoralis ahupuaa of Halepuaa Forest Reserve in the Puna District Found growing on the side of the old railroad. Preserved_Specimen ESP Field Crew Hawaii BISH 1979-08-15
449 Verbena litoralis East Maui, Haleakala Crater, Kaupo Gap Dry volcanic gravel by trail. Preserved_Specimen Flws lavender-blue. St.John, H. 21182 Maui BISH 1945-08-27
450 Verbena litoralis Waimea Audubon Center, Waimea Valley, Oahu, Hawaii Growing at the water fall just at edge of pond and edge of stream, dry to moist conditions. Preserved_Specimen Shrub, spindly, flowers purple, on spikes. Crago, L.M. 2005207 Oahu BISH 2005-10-18
451 Verbena litoralis Kipapa Gulch, Wahiawa moderately moist pineapple field Preserved_Specimen Hosaka, E.Y. 422 Oahu BISH 1931-06-18
452 Verbena litoralis Waiapaa Gulch, flat area near the road above Growing near Grevillea robusta, Psidium guajava, Molasses grass and other introduced and native plants. Preserved_Specimen Funk, E.J. 920004 Lanai BISH 1992-01-26
453 Verbena litoralis North Kohala District, Kalala ahupua'a, Lahikiola cinder cone In Kikuyu grassland Preserved_Specimen Bunch, P.M. 112 Hawaii BISH 1989-03-13
454 Verbena litoralis South Kohala: Waimea, Lalamilo Range paddock; occasional. Preserved_Specimen Branching plant 1-3 ft. Hosaka, E.Y. 2122 Hawaii BISH 1938-06-01
455 Verbena litoralis Kaalualu Bay near South Point Only one individual found in waste area. Preserved_Specimen Herb up to one m in height with small purplish flowers. Whistler, W.A. 42 Hawaii BISH 1972-04-06
456 Verbena litoralis East Maui, Upper Kaupo Gap in tall Dodonaea scrubland with Eupatorium. Nearly level with rocky rubble on ground. Preserved_Specimen Flowers purple Higashino, P.K. 885 Maui BISH 1975-06-22
457 Verbena litoralis Located on the Mauna Loa Strip road about 2.2 miles from intersection with Kona-Hilo Highway; or up Mauna Loa Strip Road about 200 meters from parking circle at Kipuka Puaulu. Plant was about 30 meters 20 deg. north of road. Plot 19 Preserved_Specimen Shrub about 4 ft. tall Newell, C.L. 966 Hawaii BISH 1967-06-23
458 Verbena litoralis near Koko Head, Kaluanui Ridge,Koolau Range guava zone; at summit Preserved_Specimen Egler, F.E. 3783 Oahu BISH 1937-07-01
459 Verbena litoralis Kaimuki, Honolulu roadside Preserved_Specimen Fosberg, F.R. 8856 Oahu BISH 1932-09-24
460 Verbena litoralis 0.5 mi NW of Puu Keekee, just SE of Parker Ranch fenceline; on rockpile, vegetation Myoporum-Chamaesyce dominant, Preserved_Specimen Wagner, W.L. 4688 Hawaii BISH 1982-11-17
461 Verbena litoralis Growing next to Kaluanui Stream Approximately 1/2 mile from ocean. Preserved_Specimen Corn, C. Oahu BISH 1978-05-18
462 Verbena litoralis Waianae Mts., Makaha Valley on trail Preserved_Specimen Hasegawa, M. Oahu BISH 1932-04-03
463 Verbena litoralis Lawai Valley; P.T.B.G Preserved_Specimen Herb w/pale blue flowers. Uncommon in valley. Herbst, D.R. 2296 Kauai BISH 1972-01-04
464 Verbena litoralis Koolau Mts.; Honolulu Forest Reserve, N of Honolulu. Manoa Cliffs Trail near Hawaiian Telephone Company facility Preserved_Specimen Herb 1 m tall; flws light violet. Lammers, T.G. 5313 Oahu BISH 1983-08-27
465 Verbena litoralis Kalaoa, HI 96740, USA OBSERVATION Curtis Daehler Hawaii HAPI 10/18/2007
466 Verbena litoralis Honolulu, Hawaii, United States OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
467 Verbena litoralis Growing next to Kaluanui Stream OBSERVATION Curtis Daehler Oahu HAPI 3/21/2007
468 Verbena litoralis Waimanalo; junction of Kalanianaole Hwy. & Hihimanu St., Albert H. Azevedo Field Roadside. Preserved_Specimen Wagner, W.L. 4826 Oahu BISH 1983-04-04
469 Verbena litoralis S half of island Ponds Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
470 Verbena litoralis Maui Preserved_Specimen Flower lavender Handy, E.S. 3320 Maui BISH 1933-06-10
471 Verbena litoralis Puuwaawaa; 4WD access track to 1986 burn (from Ranch HQ) Common roadside weed. Preserved_Specimen Takeuchi, W.N. 5752 Hawaii BISH 1989-05-01
472 Verbena litoralis East Maui, floor of Haleakala Crater 1/2 mile East of Holua Cabin along trail Preserved_Specimen Henrickson, J.S. 3939 Maui BISH 1969-07-20
473 Verbena litoralis ahupuaa of Halepuaa in the Puna District, in the Nanawale Estates subdivision The plant was growing in a fallow field at the w end of Hapu`u Road. The soil in the area is stoney being composed mostly of crushed `a`a. The area is exposed to direct sunlight and has no canopy. The dominant plants in the area area Pluchea odorata and Buddleja asiatica. Preserved_Specimen Yoshida, L. 79.053 Hawaii BISH 1979-02-28
474 Verbena litoralis Haena Preserved_Specimen Kauai BISH 1953-10-30
475 Verbena litoralis Keauhou Ranch, Kau District OBSERVATION Curtis Daehler Hawaii HAPI 3/16/2007
476 Verbena litoralis Weed of pineapple field near northern junction of Highways 37 and 377 Preserved_Specimen Much branched herb to 3 ft. tall; stem and leaves dark green, flowers purple. Herbst, D.R. 795 Maui BISH 1967-12-18
477 Verbena litoralis Kanehoa, Waianaeuka near top of ridge Preserved_Specimen Hume, E.P. 123 Oahu BISH 1931-03-22
478 Verbena litoralis East Maui, Hana District, Hana Ranch pastureland mauka of Hana Ranch Store Open pastureland with Sporobolus africanus, Digitaria ciliaris, Eragrostis pectinacea, Mimosa pudica, Sida rhombifolia, Chamaesyce prostrata, Elephantopus spicatus. Preserved_Specimen Imada, C. 200292 Maui BISH 2002-11-19
479 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Herb, woody base,1.25 m high Christophersen, E. Oahu BISH 1924-12-07
480 Verbena litoralis Kamakou Preserve (Nature Conservancy); near start of Hanalilolilo Trail Preserved_Specimen Wagner, W.L. 5398 Molokai BISH 1984-07-17
481 Verbena litoralis Wahiawa Preserved_Specimen Erect herb, 1m; flower purple Swarbrick, J.T. 0.62 Oahu BISH 1976-01-07
482 Verbena litoralis Pohakuloa dry area Preserved_Specimen Long, C.R. 4309 Hawaii BISH 1966-12-23
483 Verbena litoralis Hawaii County, HI, USA OBSERVATION Curtis Daehler Hawaii HAPI 3/25/2007
484 Verbena litoralis Maui Preserved_Specimen Forbes, C.N. 2036 Maui BISH 1920-04-19
485 Verbena litoralis Maui Preserved_Specimen Wagner, W.L. 4783 Maui BISH 1983-01-14
486 Verbena litoralis Oahu Preserved_Specimen Rémy, J. 406 Oahu BISH 1851-01-01
487 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 141 Lanai BISH 1913-10-10
488 Verbena litoralis Crater of Haleakala Preserved_Specimen Forbes, C.N. 1083 Maui BISH 1919-08-09
489 Verbena litoralis Area within 300 m west of LZ-1. UTM coordinates 752400-2275900 Preserved_Specimen Warren, S. 14 Kahoolawe BISH 1987-04-09
490 Verbena litoralis Waianae Range west of Schofield Barracks, in Kaluaa Gulch & ridge above Preserved_Specimen Sohmer, S.H. 5995 Oahu BISH 1969-08-27
491 Verbena litoralis Lanai Preserved_Specimen Munro, G.C. 292 Lanai BISH
492 Verbena litoralis Slopes of Mauna Kea (Wailei) Preserved_Specimen Forbes, C.N. 465 Hawaii BISH 1911-08-01
493 Verbena litoralis Kaua Trail, Waianae Mts Preserved_Specimen Woolford, H.A. 200 Oahu BISH 1955-12-11
494 Verbena litoralis Kealia trail Preserved_Specimen Bennett, T. 244 Oahu BISH 1982-03-25
495 Verbena litoralis Kualoa Pasture Preserved_Specimen Rock, J.F.C. 3007 Oahu BISH 1942-01-01
496 Verbena litoralis Kaholuamanoa, above Waimea Preserved_Specimen Heller, A.A. 2046 Kauai BISH 1895-09-02
497 Verbena litoralis Pauoa Valley Preserved_Specimen Garber, D.W. 339 Oahu BISH 1920-03-21
498 Verbena litoralis Honolulu, Aihualama Preserved_Specimen Russell, P. 253 Oahu BISH 1932-07-08
499 Verbena litoralis Koele Preserved_Specimen Forbes, C.N. 188 Lanai BISH 1918-06-01
500 Verbena litoralis Tantalus Rd Preserved_Specimen Forbes, A. Oahu BISH 1930-07-18
501 Verbena litoralis Waikolu Valley, head of and E. rim Preserved_Specimen Kondo, Y. Molokai BISH 1962-06-18
502 Verbena litoralis Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
503 Verbena litoralis Puu Uau, Aiea Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
504 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
505 Verbena litoralis Waianae Mts., South end Preserved_Specimen Harley, K. Oahu BISH 1956-05-04
506 Verbena litoralis Pohakaloa Burn. 1979 burn above Pohakaloa Military Camp; Mauna Kea Preserved_Specimen Adee, K.T. Hawaii BISH
507 Verbena litoralis Ko'olau Mountain; Pupukea, Paumaiu area, summit trail extending from Pupukea Road Preserved_Specimen Ziegler, M.F. Oahu BISH 1986-11-08
508 Verbena litoralis Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
509 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
510 Verbena litoralis Mokuleia Forest Reserve, Waianae Mts Preserved_Specimen Onouye, R. Oahu BISH 1934-02-04
511 Verbena litoralis Valley above country club, Honolulu Preserved_Specimen Forbes, C.N. Oahu BISH 1908-07-08
512 Verbena litoralis S half of island Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
513 Verbena litoralis Palolo Valley, middle ridge forest Preserved_Specimen Christophersen, E. 1666 Oahu BISH 1931-03-26
514 Verbena litoralis Palaau, west side of airport pineapple field; dry, exposed, windy. Preserved_Specimen Herb to sub-shrub with blue flowers Herbst, D.R. 5092 Molokai BISH 1974-11-11
515 Verbena litoralis Waipio Preserved_Specimen Swanson, R. Hawaii BISH 1931-08-29
516 Verbena litoralis Maui Preserved_Specimen Neal, M.C. Maui BISH 1933-08-01
517 Verbena litoralis unknown Preserved_Specimen Mann, H. 437 BISH 1864-01-01
518 Verbena litoralis Kilauea, Kaloko Reservoir Preserved_Specimen Forbes, C.N. 572 Kauai BISH 1916-10-11
519 Verbena litoralis Palawai Preserved_Specimen Munro, G.C. 365 Lanai BISH 1928-08-01
520 Verbena litoralis Mt. Kalena, Waianae Lantana grove Preserved_Specimen herb 4 ft. Grant, M.L. 7445 Oahu BISH 1934-09-30
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