Description
Key Characters:
Growth Form:
Gynodioecious shrubs, 3–8 dm tall.
Stems:
Stems often swollen at the nodes, terete (cylindrical), 4–angled or –ridged, erect or strongly ascending, glabrous, internodes 0.5–4.1 cm long, green to greenish purple, terete (cylindrical), 4–angled or –ridged, lateral branches of primary stems usually elongating several or more nodes below the inflorescence.
Roots:
Fibrous root system.
Leaves:
Leaves simple.
Opposite.
Blades elliptic to elliptic–oblanceolate or oblanceolate, 1.5–3 (–6) cm long, 0.4–1.4 (–2.2) cm wide.
Apex acute to acuminate.
Base leaf bases often connate around the stem, base cuneate.
Surfaces yellowish green; blades weakly coriaceous.
Margins entire, slightly thickened and weakly revolute.
Only the midvein evident.
Petioles 0.3–0.8 cm long.
Stipules absent.
Flowers:
Flowers in inflorescence terminal, laterally contracted, somewhat congested; the hermaphroditic inflorescences 3.1–12.3 cm long and with 22–170 flowers, less congested and broader than the pistillate inflorescences, these 2.3–11.4 cm long and with 30–140 flowers; bracts of central axis 2–15 mm long, those of branches and flowers 1–1.8 mm long, green to purple, linear to subulate.
Flowers bisexual (perfect) or pistillate, (4–)5-merous.
Calyx of hermaphroditic flowers with sepals 3.4–4.5 mm long, narrowly ovate, broadly and shallowly navicular, oriented at 60° to 90° angle to the pedicel, green or tinged purple toward the apex, opaque, reflexed and convex at the base, producing a small transverse bulge, glabrous, margins scarious, sometimes somewhat irregularly undulate, entire, glabrous, apex obtuse. Pistillate flowers: sepals 2.6–3.2 mm long, narrowly ovate, green or yellowish green, pale yellow at the base, opaque, venation pale and translucent, reflexed and convex at the base, producing a small transverse bulge, broadly navicular, oriented at 60° to 100° angle to the pedicel, margins scarious and often irregularly undulate, entire or with 1–several lacerations, glabrous, apex often inrolled, obtuse.
Corolla (petals) absent.
Stamens 10 (hermaphroditic flowers), the filaments slightly dimorphic, the antisepalous whorl 4.9–5.7 mm long, the alternate whorl 4.5–5.3 mm long; anthers 0.8–1.2 mm long, yellow. Pistillate flowers: Stamens 10, vestigial; filaments unequal, the antisepalous whorl 0.5–1.75 mm long, the alternate whorl 0.4–0.65 mm long; anthers 0.2–0.4 mm long, white, not producing pollen.
Ovary superior; styles 3–4 (–5) (pistillate flowers), or 3–4 (hermaphroditic flowers).
Fruit:
Capsules 3.5–5 mm long; narrowly ovoid.
Seeds relatively few; 0.8–0.9 mm long; reniform; compressed; the surface papillose; papillae of the margins larger than those of the faces; perisperm starchy; hard or rarely soft; true endosperm absent.
Ploidy:
2n = 52*–56*; 60*
Habitat:
Known only from steep; dry slopes in largely alien shrubland.
Elevation Range:
ca. 125 m.