Geniostoma hedyosmifolium

(Baill.) Byng & Christenh. (2018)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Gentianales Family: Loganiaceae Genus: Geniostoma

kāmakahala [kamakahala]

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Description

Key Characters:

Growth Form: Terrestrial or sometimes epiphytic, sprawling, liana-like shrubs, erect, branched shrubs, or sometimes shrubby trees 0.4–4.5 m tall.

Stems: Stems angled or slightly winged, glabrous or the uppermost moderately or sometimes densely short–hispidulous.

Roots:

Leaves: Leaves simple. Opposite. Blades usually narrowly to broadly elliptic or oblanceolate, sometimes obovate, (3.5–)5–13.5 cm long, 1.8–5.5(–6) cm wide. Apex abruptly short-acuminate or acuminate. Base cuneate to broadly cuneate or sometimes rounded. Upper surfaces glabrous; lower surfaces glabrous or sparsely to sometimes moderately short–hispid, especially or exclusively on veins; leaves green on both surfaces or lower surfaces pale green. Margins flat or slightly revolute, at least when dry. Upper surface with veins slightly impressed, lower surface with veins not or moderately raised. Petioles 0–1.5(–2.2) cm long. Stipules interpetiolar, may also be intrapetiolar, membranous, connate into a cup 3–6 mm long, not or slightly adnate to base of petioles, glabrous, quickly splitting, often with thickened pale brown margins, eventually deciduous.

Flowers: Flowers in inflorescences terminal, flowers (2–)5–25(–30) in sessile or subsessile, simple or compound, contracted or relatively open cymes, glabrous to moderately or densely short–hispidulous, peduncles 0–3 mm long, pedicels 3–7 mm long, elongating to 5–18 mm long in fruit, bracts linear–filiform or sometimes narrowly elliptic, 6–9 mm long, bracteoles linear–filiform to filiform, 6–8 mm long, ± with a few scattered hairs. Flowers bisexual (perfect) or functionally unisexual, actinomorphic. Calyx lobes 5–6, unequal to subequal, united at base, ± imbricate, oblong–lanceolate to linear–lanceolate, sometimes narrowly elliptic with a constricted base, (4.5–)7–16 mm long, 1.6–3(–6) mm wide, perhaps enlarged only in fruit, conspicuously 5–7–nerved, glabrous or sometimes sparsely short–hispidulous, margins hyaline, glabrous or ciliate, apex acute to attenuate, inner surface of Calyx often with colleters at the base, margin mostly fimbriate. Corolla 5-merous, campanulate to rotate, orangish yellow, 11.3–22 mm long, the tube flaring toward mouth, 6.8–15 mm long, glabrous or sometimes sparsely short–hirsute, the lobes 5–6, narrowly deltate, 4.5–7.3 mm long, apex acute, moderately white hirsute on the tube within and on lower part of lobes, lobes imbricate or contorted in bud. Stamens 5 alternipetalous, attached to mouth of corolla, exserted, usually recurved after anthesis, glabrous or hairy; filaments mostly short; anthers 2-celled, introrse, connective often extended to form an apical appendage, opening by longitudinal slits. Ovary superior or partly inferior, carpels hairy or glabrous, 2– or 3-locular, sometimes partitions imperfect in upper part of ovary; ovules numerous, amphitropous, placentation axile; style terminal, mostly short; stigma clavate, ellipsoid, to globular, sometimes ± oblong, about as large as ovary.

Fruit: Capsules ovoid–ellipsoid to ellipsoid or sometimes broadly ovoid; 12–20 mm long; usually sparsely hirtellous; 2–3(4)–valved; valves usually conspicuously keeled in upper part; at least while maturing; keel 1.5–3.5 mm wide; apex with a beak 3.5–7 mm long; septicidal to sub–septifragal; maturing to green; with or without a black–purple tinge; usually becoming brown–black when over–mature. Seeds ca. 1 mm long; ± winged at both ends or all around; embedded in juicy orange to red pulp; numerous; ellipsoid to subglobular; intruded on the hilar side; surface mostly minutely papillose to areolate; brown to black; endosperm thick; fleshy; oily; proteinaceous; with saccharose or hemicellulose; embryo small; straight.

Ploidy:

Habitat: Scattered or locally common primarily in wet forest but extending to mesic forest or sometimes bog margins.

Elevation Range: 700–1,590 m.

Historical Distribution

Images

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Molokai Endemic
Lana'i Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Global Fl. 4: 145 (2018)

Other References

Wagner et al. 1990:856 (Mo, L, M, H [as Labordia hedyosmifolia]); Gibbons et al. 2012/Christenhusz et al. 2018:145 (Syn. L. hedyosmifolia = G. hedyosmifolium)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Geniostoma hedyosmifolium W Maui, Lahaina District, Honokohau, Nakalalua Growing in dense shade in Wet Forest. Preserved_Specimen Oppenheimer, H.L. 60009 Maui BISH 2000-06-02
2 Geniostoma hedyosmifolium Makawao District, Honopou AHU, Tr 2(40), NW slope of Haleakala, Koolau Forest Reserve; Along Kailua Stream Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 2776 Maui BISH 1980-07-10
3 Geniostoma hedyosmifolium Kona-Hilo Hwy, ca. 6.8 miles toward Hilo from Hongo Store Metrosideros-Cibotium forest community Preserved_Specimen Newell, C.L. 1003 Hawaii BISH 1967-06-26
4 Geniostoma hedyosmifolium Piihonua; Hilo Forest Reserve, Site 26 OS Primary vegetation: Metrosideros polymorpha (pubescent), Ilex anomala, Myrsine lessertiana, Cheirodendron trigynum & Coprosma ochracea. Preserved_Specimen Adee, K.T. Hawaii BISH 1978-10-26
5 Geniostoma hedyosmifolium East Maui, Hana District, Doolau FR, west fork of Heleleikeoha stream Along perennial stream. Vegetation: Metrosideros, Cheirodendron, Broussaisia, Dicranopteris, Melicope, Sadleria, Cibotium, Clermontia, Cyanea, Astelia, Peperomia, Athyrium, Kadua, Lycopodiella, Vaccinium, Labordia, Isachne, Eragrostis, Deschampsia, Pipturus, Pilea, Machaerina, Dryopteris, Coprosmia, Psychotria, Cyrtandra, Prunella, Ageratina, Tibouchina, Plantago major, Rubus hawaiensis, Korthalsella. Preserved_Specimen Oppenheimer, H.L. 51419 Maui BISH 2014-05-28
6 Geniostoma hedyosmifolium South Kona District; Kipahoehoe AHU Tr 68(24); West slopes of Mauna Loa; 0.5-1 km south of Kaapuna 1950 lava flow Mesic `ohi`a forest with other native trees and a native shrub, matted-fern understory. Growing on `a`a with little soil. Growing on `a`a. Preserved_Specimen Warshauer, F.R. 2006 Hawaii BISH 1978-07-22
7 Geniostoma hedyosmifolium East Maui, Hana District, Waihoi Valley, Puu Hoolio Under Cheirodendron, in Dicranopteris & Paspalum conjugatum. Preserved_Specimen Harrison, B. 2520 Maui BISH 1972-07-20
8 Geniostoma hedyosmifolium Puna District, 26 miles up Highway 11, Mauna Loa Estates, corner of Ruby and 9th Closed Metrosideros forest with Cibotium dominated understory. Preserved_Specimen Davis, J. 367 Hawaii BISH 1980-09-14
9 Geniostoma hedyosmifolium Kilauea Iki Crater East wall of crater On moist wooded part. Preserved_Specimen Fosberg, F.R. 46049 Hawaii BISH 1965-03-19
10 Geniostoma hedyosmifolium South Kohala District; Awini AHU. Tr 78(11); Kohala Mountains; 0.5 km northeast of Honokanenui Stream Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 2482 Hawaii BISH 1979-08-01
11 Geniostoma hedyosmifolium Upper Hamakua Ditch, Kohala Mountains Over lava tube. Preserved_Specimen Corn, C. Hawaii BISH 1984-03-30
12 Geniostoma hedyosmifolium Wailuku District, Kahakuloa AHU, Tr 38(9), West Maui Mountains, in bog between Makamakaole & Huluhulupueo Streams Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 3131 Maui BISH 1980-08-25
13 Geniostoma hedyosmifolium North Kona District; Keauhou AHU Tr 57(160); northwest slope of Mauna Loa, east of Mamalahoe Highway Mesic `ohi`a forest with other native trees and a native shrub, matted-fern understory. Growing on `a`a with little soil. Growing on `a`a. Preserved_Specimen Warshauer, F.R. 1862 Hawaii BISH 1978-06-18
14 Geniostoma hedyosmifolium Waiakea; Site 27 OS Kulani Project Closed healthy forest. Soil a shallow, well drained drained organic over an a`a flow. Primary vegetation: Metrosideros polymorpha (pubescent), Ilex anomala, Myrsine lessertiana, Myrsine sandwicensis, Cheirodendron trigynum & Coprosma spp. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-22
15 Geniostoma hedyosmifolium Along Papaala Pali Rain forest Preserved_Specimen Degener, O. 10274 Molokai BISH 1928-06-06
16 Geniostoma hedyosmifolium South Kona District; Kealakekua AHU Tr 60(119/120); W slope of Mauna Loa, 2-4 km east of Keopuka Wet `ohi`a forest with other native trees and a treefern and native shrub understory. Growing on `a`a lava. Preserved_Specimen Jacobi, J.D. 1141 Hawaii BISH 1978-08-03
17 Geniostoma hedyosmifolium South of 26 Miles, Puna in jungle. Preserved_Specimen Degener, O. 33612 Hawaii BISH 1968-11-09
18 Geniostoma hedyosmifolium Piihonua; Hilo Forest Reserve, Site 26 OS Pubescent ohia dieback. Soil an extremely shallow organic found over an a`a flow. Primary vegetation: Metrosideros polymorpha (pubescent), Ilex anomala, Myrsine lessertiana, Cheirodendron trigynum & Coprosma ochracea. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-27
19 Geniostoma hedyosmifolium W Maui, Lahaina District, Haelaau-Puu Kukui Trail Rain forest Preserved_Specimen Ewart III, G.R. 114 Maui BISH 1928-12-18
20 Geniostoma hedyosmifolium Upper Olaa Forest Reserve On hapu`u. Growing with Cibotium and Pneumatopteris on a steep slope. Scattered tall Metrosideros with closed Cibotium understory. Preserved_Specimen Davis, J. 407 Hawaii BISH 1980-10-22
21 Geniostoma hedyosmifolium Waiakea; Upper Waiakea Forest Reserve, Site 15 OS Ohia-treefern dieback. Soil a deep and moderately well drained organic/ash; substrate unknown. Primary vegetation: Metrosideros polymorpha, Coprosma spp., Ilex anomala, Gouldia sp., Cheirodendron trigynum & Psychotria sandwicensis. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-07
22 Geniostoma hedyosmifolium Olaa; Site 31 OS Olaa Forest Reserve Closed healthy forest. Soil a moderately deep & well drained organic/ash over a pahoehoe flow. Primary vegetation: Metrosideros polymorpha, Ilex anomala, Myrsine lessertiana, Cheirodendron trigynum, Coprosma spp. & Psychotria hawaiiensis. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-28
23 Geniostoma hedyosmifolium Kona-Hilo Hwy, ca. 6.8 miles toward Hilo from Hongo Store Metrosideros-Cibotium forest community Preserved_Specimen Newell, C.L. 657 Hawaii BISH 1966-08-28
24 Geniostoma hedyosmifolium Kona-Hilo Highway. Ca. 6.8 miles toward Hilo from Hongo Store Metrosideros-Cibotium forest community. Preserved_Specimen Newell, C.L. 658 Hawaii BISH 1966-08-28
25 Geniostoma hedyosmifolium East Maui, Hana District, Tr 17(46), Along Kaukauai Gulch, SW kipahulu Valley near waterfall.i Ridge Wet koa-`ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 2861 Maui BISH 1980-07-22
26 Geniostoma hedyosmifolium Kona-Hilo Hwy, ca. 6.8 miles toward Hilo from Hongo Store Metrosideros-Cibotium forest community Preserved_Specimen Newell, C.L. 656 Hawaii BISH 1966-08-28
27 Geniostoma hedyosmifolium W Maui, Puu Kukui On ridge In rain forest Preserved_Specimen Neal, M.C. Maui BISH 1933-08-16
28 Geniostoma hedyosmifolium South Kona District; Kealakekua AHU Tr 60(119/120); W slope of Mauna Loa, 2-4 km east of Keopuka Wet `ohi`a forest with other native trees and a treefern and native shrub understory. Growing on `a`a lava. Preserved_Specimen Jacobi, J.D. 1141 Hawaii BISH 1978-08-03
29 Geniostoma hedyosmifolium East Maui, Hana District, Waihoi Valley, N fork of Waiohonu Along bank with Dicranopteris. Preserved_Specimen Harrison, B. 2626 Maui BISH 1972-07-23
30 Geniostoma hedyosmifolium W Maui, near top of Mount Eke Open windswept rain forest. Preserved_Specimen Degener, O. 10259 Maui BISH 1927-08-30
31 Geniostoma hedyosmifolium Kawela AHU. Tr 5(5); Top of Papaala Pali, Kaunuohua Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Stemmermann, R.L. 3915 Molokai BISH 1979-06-21
32 Geniostoma hedyosmifolium W Maui, Lahaina District, Haelaau Lower rain forest Preserved_Specimen Bryan Jr., E.H. 639 Maui BISH 1928-12-20
33 Geniostoma hedyosmifolium Olaa Forest Reserve. On road from Hilo to Kulani Prison Dense, swampy Metrosideros-Cibotium montane rain forest. Pritchardia & Joinvillea present. Preserved_Specimen Hatheway, W.H. 442 Hawaii BISH 1950-10-31
34 Geniostoma hedyosmifolium East Maui, Waihoi Valley, Unnamed stream at base of N wall of valley On bank with Vaccinium Preserved_Specimen Harrison, B. 2571 Maui BISH 1972-07-21
35 Geniostoma hedyosmifolium Piihonua; Hilo Forest Reserve, Site 05 OS Pubescent ohia dieback. Soil an extremely shallow & excessively drained organics over an a`a flow. Primary vegetation: Metrosideros polymorpha (pubescent), Ilex anomala, Myrsine lessertiana, M. sandwicensis & Cheirodendron trigynum. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-25
36 Geniostoma hedyosmifolium Site 24 OS; Olaa, Olaa Forest Reserve Closed healthy forest; assoc. spp.: Metrosideros polymorpha, Ilex anomala, Myrsine lessertiana, Cheirodendron trigynum, Coprosma spp., Psychotria hawaiiensis & Gouldia sp. Soil a moderately deep and well drained organic/ash found over an a`a flow. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-07-09
37 Geniostoma hedyosmifolium South Kona District, Kealakekua Ranch. Kelalakekua ahupuaa, road to Papaloa Mesic Metrosideros/Cibotium forest with Freycinetia common. Preserved_Specimen Wagner, W.L. 5945 Hawaii BISH 1988-03-11
38 Geniostoma hedyosmifolium Up Kulani Prison Road, 5.3 miles toward Prison from intersection with Volcano Road Metrosideros-Cibotium forest community. Preserved_Specimen Newell, C.L. 695 Hawaii BISH 1966-09-01
39 Geniostoma hedyosmifolium North Hilo District; Umauma AHU. Tr 18(100); east slope of Mauna Kea on ridge between Umauma Stream & Nauhi Gulch Wet koa-`ohi`a forest with other native trees, a treefern & native shrub understory. Preserved_Specimen Warshauer, F.R. 1342 Hawaii BISH 1977-08-21
40 Geniostoma hedyosmifolium Piihonua; Hilo Forest Reserve, Site 05 OS Pubescent ohia dieback. Soil an extremely shallow & excessively drained organics over an a`a flow. Primary vegetation: Metrosideros polymorpha (pubescent), Ilex anomala, Myrsine lessertiana, M. sandwicensis & Cheirodendron trigynum. Preserved_Specimen Adee, K.T. Hawaii BISH
41 Geniostoma hedyosmifolium Kau Forest Resesrve, Kau District Open Metrosideros - Acacia koa forest with closed Cibotium understory. Preserved_Specimen Davis, J. 647 Hawaii BISH 1981-11-19
42 Geniostoma hedyosmifolium Waiakea; Site 19 OS Upper Waiakea Forest Reserve Closed healthy forest. Soil a moderately shallow, well drained organic found over an a`a flow. Primary vegetation: Metrosideros polymorpha, Ilex anomala, Myrsine lessertiana, M. sandwicensis, Cheirodendron trigynum & Broussaisia arguta. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-14
43 Geniostoma hedyosmifolium South Kona District; Alika AHU Tr 69(42/43); SW slope of Mauna Loa, 0.7 km south of Alika 1919 and 3 km north of Hoopuloa 1926 flows Wet `ohi`a forest with native shrubs/matted fern and some treeferns in the understory. Growing on `a`a with little soil. Preserved_Specimen Jacobi, J.D. 1113 Hawaii BISH 1978-07-23
44 Geniostoma hedyosmifolium Upper Olaa Forest Reserve Growing with Cibotium and Pneumatopteris on a steep slope. Scattered tall Metrosideros with closed Cibotium understory. Preserved_Specimen Davis, J. 407 Hawaii BISH 1980-10-22
45 Geniostoma hedyosmifolium East Maui, Hana District, Keanae AHU, Tr 6(31), N slope of Haleakala, Koolau Forest Reserve, ridge along E Wailuaiki Stream Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 2602 Maui BISH 1980-06-01
46 Geniostoma hedyosmifolium W Maui, Lahaina District, Lihau Montane Wet Shrubland. A sprawling groundcover with Metrosideros, Cheirodendron, Coprosma, Broussaisia, Psychotria, Sadleria, Liparis, Stenogyne and Peperomia. Preserved_Specimen Welton, P. 1079 Maui BISH 1991-07-13
47 Geniostoma hedyosmifolium Kulani Project, S. Hilo District In closed Metrosideros forest with Gouldia, Styphelia, Alyxia, Cibotium glaucum, and Stenogyne calaminthoides. `A`a substrate densely moss covered. Preserved_Specimen Davis, J. 455 Hawaii BISH 1981-02-20
48 Geniostoma hedyosmifolium W Maui, Puu Kukui On ridge In rain forest Preserved_Specimen Neal, M.C. Maui BISH 1933-08-16
49 Geniostoma hedyosmifolium head of Waikolu Valley Rain-forest Preserved_Specimen Degener, O. 22167 Molokai BISH 1952-03-15
50 Geniostoma hedyosmifolium Wailau AHU. Tr 20(6); Between Wailau & Pelekunu Valleys; east ridge of Olokui Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 2958 Molokai BISH 1980-08-03
51 Geniostoma hedyosmifolium South Kona District; Puu Kapu AHU Tr 80(9); Kohala Mountains; northeast of Puu Kaiholena Wet `ohi`a forest with other native trees and a treefern and native shrub understory. Preserved_Specimen Stemmermann, R.L. 3986 Hawaii BISH 1979-08-01
52 Geniostoma hedyosmifolium W Maui, Lahaina District, Puu Kukui trail, between Haelaau cabin and Nakalalu In Metrosideros-Dicranopteris montane wet forest with Clermontia, Pittosporum, Stenogyne, Ilex, Broussaisia, Coprosma, Labordia, Cyrtandra, Myrsine, Athyrium sandwichianum, Tibouchina herbacea. Preserved_Specimen Perlman, S. 6860 Maui BISH 1987-11-14
53 Geniostoma hedyosmifolium South Hilo District; Makahanaloa AHU. Tr 20(37); east slope of Mauna Kea, along Kapue Stream; Hilo Forest Reserve Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 1469 Hawaii BISH 1977-10-05
54 Geniostoma hedyosmifolium South Kona District; Kipahoehoe AHU Tr 68(23); West slopes of Mauna Loa; 0.5-1 km south of Kaapuna 1950 lava flow Mesic `ohi`a forest with other native trees and a native shrub, matted-fern understory. Growing on `a`a with little soil. Growing on `a`a. Preserved_Specimen Warshauer, F.R. 2005 Hawaii BISH 1978-07-22
55 Geniostoma hedyosmifolium Kulani Prison Road, 7 miles east of Kulani Shaded rain forest of Metrosideros, Cibotium, Coprosma, Cyrtandra, Antidesma, etc. Preserved_Specimen Henrickson, J.S. 4111 Hawaii BISH 1969-08-07
56 Geniostoma hedyosmifolium South Hilo District; Piihonua AHU. Tr 22(66); east slope of Mauna Kea on ridge south of Nukupahu Gulch Wet `ohi`a forest with other native trees, a treefern & native shrub understory. Preserved_Specimen Warshauer, F.R. 1630 Hawaii BISH 1977-12-16
57 Geniostoma hedyosmifolium East Maui, Hana District, Keanae AHU, Tr 4(38), N slope of Haleakala, Keanae Valley Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Growing on `a`a substrate. Preserved_Specimen Warshauer, F.R. 2821 Maui BISH 1980-07-14
58 Geniostoma hedyosmifolium W Maui, Lahaina District, headwaters of Honokohau at Kahoolewa Ridge In wet forest w. Metrosideros, Cheirodendron, Ilex, Myrsine, Melicope, Clermontia, Cyrtandra, Smilax, Astelia, Athyrium, Sadleria, Cibotium, Dryopteris. Preserved_Specimen Oppenheimer, H.L. 50228 Maui BISH 2002-05-20
59 Geniostoma hedyosmifolium Kainalu Stream Vegetation:: Metrosideros, Cheirodendron, Dicranopteris Diplopterygium, Broussaisia, Elaphoglossum, Cibotium, Sadleria, Cyrtandra, Coprosma, Kadua, Labordia, Clermontia, Lobelia, Peperomia, Tibouchina, Clidemia. Preserved_Specimen Oppenheimer, H.L. 60907 Molokai BISH 2009-06-04
60 Geniostoma hedyosmifolium Olinda, Pipe-line trail Dense, dark rain forest. Preserved_Specimen Degener, O. 10264 Maui BISH 1927-06-21
61 Geniostoma hedyosmifolium Kulani Prison Road, 7 miles east of Kulani Shaded rain forest of Metrosideros, Cibotium, Coprosma, Cyrtandra, Antidesma, etc. Preserved_Specimen Henrickson, J.S. 4111 Hawaii BISH 1969-08-07
62 Geniostoma hedyosmifolium east Kopiliula 50% open 7 m canopy; Acacia, Metrosideros, uluhe, ohia, Cheirodendron, Ilex, Clermontia arborescens, Scaevola chamissoniana, Myrsine lessertiana, Melicope, Hedyotis, Coprosma, Perrottetia, Broussaisia, Vaccinium, Styphelia, Cyrtandra, Rubus hawaiensis, ferns, bryophytes Preserved_Specimen Wood, K.R. 11316 Maui BISH 2005-05-04
63 Geniostoma hedyosmifolium South Hilo District; Paukaa AHU. Tr 21(73); east slope of Mauna Kea, between Awehi & Honolii Streams Wet matted fern and/or native shrub community with scattered `ohi`a and other native trees. Preserved_Specimen Warshauer, F.R. 1399 Hawaii BISH 1977-09-12
64 Geniostoma hedyosmifolium Up Kulani Prison Road, 5.3 miles toward Prison from intersection with Volcano Road Metrosideros-Cibotium forest community Preserved_Specimen Newell, C.L. 693 Hawaii BISH 1966-09-01
65 Geniostoma hedyosmifolium East Maui, Mountains above Hana Swamp. Preserved_Specimen Forbes, C.N. 2680 Maui BISH 1920-07-05
66 Geniostoma hedyosmifolium Kawela AHU Tr 5(4); Along Papaala Pali, Kaunuohua; pit crater near S(4) Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Stemmermann, R.L. 3938 Molokai BISH 1979-06-21
67 Geniostoma hedyosmifolium East Maui, Hana District, Keanae AHU, Tr 4(45), N slope of Haleakala, lower Keanae Valley Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Growing on a`a substrate. Preserved_Specimen Warshauer, F.R. 2829 Maui BISH 1980-07-15
68 Geniostoma hedyosmifolium Hanalilolilo Trail, below Pepeopae Bog In dense rain forest. Preserved_Specimen Wagner, W.L. 5414 Molokai BISH 1984-07-17
69 Geniostoma hedyosmifolium Keauhou Ramch, Kau District, near jeep trail connecting Kilauea Forest Reserve boundary and powerline trail Tall closed Metrosideros forest in kipuka surrounded by more recent lava. Preserved_Specimen Cuddihy, L.W. 48 Hawaii BISH 1979-12-18
70 Geniostoma hedyosmifolium Wailau AHU. Tr 19(25/26; Between Wailau & Pelekunu Valleys; along Kapapa Pali Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 2968 Molokai BISH 1980-08-04
71 Geniostoma hedyosmifolium Koolau Ahupuaa, East Maui, downslope of Puualaea and near stream bank of Hanawi Stream west fork. Transect MT, Station 7. Wet ohia forest with other native trees and a native shrub-matted fern understory. Preserved_Specimen Warshauer, F.R. 2539 Maui BISH 1980-05-20
72 Geniostoma hedyosmifolium Kilauea, Hilo side of National Park Wet tree fern forest. Preserved_Specimen Fosberg, V.O. 54 Hawaii BISH 1936-08-10
73 Geniostoma hedyosmifolium Munro Trail, near top. Shrubby rain forest near top. Preserved_Specimen Degener, O. 24177 Lanai BISH 1957-08-13
74 Geniostoma hedyosmifolium Makawao District, Haiku Uka AHU, Tr 1(11), NW slope of Haleakala, Koolau FR; on ridge west of Opana Gulch Wet koa-`ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 2755 Maui BISH 1980-06-29
75 Geniostoma hedyosmifolium East Maui, Hana DistrictY. Tr 10(15), NE slope of Haleakala, Hana Forest Reserve, along Kalapawili Ridge Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Higashino, P.K. 9286 Maui BISH 1980-06-28
76 Geniostoma hedyosmifolium Piihonua; Hilo Forest Reserve, Site 04 OS Stunted ohia wetland dieback; soil moderately deep yet have poorly drained organic surface over a thin ash deposit on pahoehoe flow. Primary vegetation: Metrosideros polymorpha, Acacia koa on well drained hummocks; Ilex anomala & Myrsine lessertiana. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-06
77 Geniostoma hedyosmifolium W Maui, Lihau Montane Wet Shrubland at 4100 ft near the summit of Lihau with Metrosideros, Clermontia arborescens, Coprosma, Vaccinium calycinum, Broussaisia, Dubautia spp. Hedyotis, native herbs, sedges and ferns Preserved_Specimen Welton, P. 1097 Maui BISH 1991-07-13
78 Geniostoma hedyosmifolium North Honaunau. First transect in North Honaunau, at Swindale's Stennis-McIntyre Plots Wet Metrosideros rain forest, heavy Freycinetia-Broussaisia tangle for undergrowth. Preserved_Specimen Herbst, D.R. 1725 Hawaii BISH 1970-07-08
79 Geniostoma hedyosmifolium East Maui, Hana District, Tr 8(41), N slope of Haleakala, just S of Kuhiwa Gulch Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 2651 Maui BISH 1980-06-15
80 Geniostoma hedyosmifolium Puna District; Olaa AHU. Tr 28(145); east slope of Mauna Loa; S of Stainback highway; Olaa Forest Reserve Wet exotic tree, `ohi`a forest with a matted-fern & native shrub understory. Preserved_Specimen Warshauer, F.R. 1280 Hawaii BISH 1977-08-04
81 Geniostoma hedyosmifolium South Kona District; Alika AHU Tr 69(42/43); SW slope of Mauna Loa, 0.7 km south of Alika 1919 and 3 km north of Hoopuloa 1926 flows Wet `ohi`a forest with native shrubs/matted fern and some treeferns in the understory. Growing on `a`a with little soil. Preserved_Specimen Jacobi, J.D. 1113 Hawaii BISH 1978-07-23
82 Geniostoma hedyosmifolium W Maui, Helu Summit Amfac Survey transect 7 In Metrosideros-Cheirodendron montane wet forest with Eurya, Clermontia, Melicope, Cyrtandra, Labordia, Stenogyne, Coprosma Preserved_Specimen Perlman, S. 10739 Maui BISH 1989-11-07
83 Geniostoma hedyosmifolium Papaikou; Hilo Forest Reserve, Site 35 OS Stunted ohia wetland dieback; soil a somewhat poorly drained organic surface over deep ash. Substrate unknown. Primary vegetation: Metrosideros polymorpha, Myrsine lessertiana, Myrsine sandwicensis, Cheirodendron trigynum & Pelea clusiifolia. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-04-03
84 Geniostoma hedyosmifolium Hana District. Kawaipapa stream, along stream on wet cliff walls, above Nahiku-Hana area Metrosideros-Cheirodendron trigynum wet forest with Sadleria spp., Pipturus albidus, Clermontia grandiflora, Dubautia plantaginea, Cyrtandra grayana, Broussaisia arguta, Coprosma foliosa, Perrottetia sandwicensis, Clermontia kakeana, Polyscias oah Preserved_Specimen Perlman, S. 23387 Maui BISH 2013-04-11
85 Geniostoma hedyosmifolium Waiakea; Site 19 OS Upper Waiakea Forest Reserve Closed healthy forest. Soil a moderately shallow, well drained organic found over an a`a flow. Primary vegetation: Metrosideros polymorpha, Ilex anomala, Myrsine lessertiana, M. sandwicensis, Cheirodendron trigynum & Broussaisia arguta. Preserved_Specimen Adee, K.T. Hawaii BISH 1978-08-30
86 Geniostoma hedyosmifolium Along Saddle Road, 19 miles from Hilo In rich, forested kipuka south of road Preserved_Specimen Degener, O. 19792 Hawaii BISH 1949-08-29
87 Geniostoma hedyosmifolium East Maui, Hana District, Tr 12(9), NE slope of Haleakala, Hana Forest Reserve, S fork of Kalapawili Gulch Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Growing on a`a substrate. Preserved_Specimen Warshauer, F.R. 2918 Maui BISH 1980-07-28
88 Geniostoma hedyosmifolium W Maui, Lahaina District, Violet Lake area of Puu Kukui In bog proper & also in margins. Associated with Metrosideros, Sadleria, Ilex anomala, Lobelia gaudichaudii gloria-montis, Argyroxiphium caliginis, A. grayanum, etc. Preserved_Specimen Gustafson, R. 2042 Maui BISH 1980-12-11
89 Geniostoma hedyosmifolium South Hilo District; Paukaa AHU. Tr 21(68); east slope of Mauna Kea, ridge between Awehi & Honolii Streams Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 1397 Hawaii BISH 1977-09-11
90 Geniostoma hedyosmifolium Kilauea. Near sulphur banks Shaded earthquake crack. Preserved_Specimen Degener, O. 10246 Hawaii BISH 1929-10-21
91 Geniostoma hedyosmifolium Ukumehame Gulch Wet Metrosideros forest. Preserved_Specimen St.John, H. 25693 Maui BISH 1955-12-28
92 Geniostoma hedyosmifolium East Maui, Hana District, Koolau FR, west fork of Heleleikeoha stream. Along perennial stream. Vegetation: Metrosideros, Cheirodendron, Broussaisia, Dicranopteris, Melicope, Sadleria, Cibotium, Clermontia, Cyanea, Astelia, Peperomia, Athyrium, Kadua, Lycopodiella, Vaccinium, Labordia, Isachne, Eragrostis, Deschampsia, Pipturus, Pilea, Machaerina, Dryopteris, Coprosmia, Psychotria, Cyrtandra, Prunella, Ageratina, Tibouchina, Plantago major, Rubus hawaiensis, Korthalsella. Preserved_Specimen Oppenheimer, H.L. 51427 Maui BISH 2014-05-29
93 Geniostoma hedyosmifolium Kalawao District; Waikolu AHU Tr 4(5/6); Along East fork of Waikolu Stream Terrestrial, semi-shade. Preserved_Specimen Jacobi, J.D. 1548 Molokai BISH 1979-07-15
94 Geniostoma hedyosmifolium East Maui, Hana District, Tr 11(9), NE slope of Haleakala, Hana Forest Reserve, Along E part of Kalapawili Ridge Wet open bog with scattered native shrubs. Preserved_Specimen Warshauer, F.R. 2908 Maui BISH 1980-07-27
95 Geniostoma hedyosmifolium South Hilo District; Waiakea AHU. Tr 26(41); east slope of Mauna Loa, across 1852 lava flow Wet `ohi`a forest with other native trees, a treefern & native shrub understory. Growing on mucky ground on pahoehoe. Preserved_Specimen Warshauer, F.R. 1614 Hawaii BISH 1977-12-10
96 Geniostoma hedyosmifolium W Maui, Lahaina District, up trail from Haelaau Cabin at transect station 4.6510 mesic forest Preserved_Specimen Annable, C.R. 3852 Maui BISH 1998-03-05
97 Geniostoma hedyosmifolium Piihonua; Hilo Forest Reserve, Site 05 OS Pubescent ohia dieback. Soil an extremely shallow & excessively drained organics over an a`a flow. Primary vegetation: Metrosideros polymorpha (pubescent), Ilex anomala, Myrsine lessertiana, M. sandwicensis & Cheirodendron trigynum. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-25
98 Geniostoma hedyosmifolium Piihonua; Hilo Forest Reserve, Site 09 OS Ohia wetland dieback. Most of the soils are shallow organic and poorly-drained over a pahoehoe flow. Some areas are well-drained hummocks. Primary vegetation: Metrosideros polymorpha, Acacia koa, Cheirodendron trigynum & Myrsine lessertiana. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-06
99 Geniostoma hedyosmifolium South Kona District; Honokau AHU Tr 65(57); W slopes of Mauna Loa; 1- 2 km north of Honokua 1950 lava flow Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Growing on pahoehoe. Preserved_Specimen Warshauer, F.R. 1934 Hawaii BISH 1978-07-02
100 Geniostoma hedyosmifolium Puna District. Hawaiian Orchis Isle Estates Occasional occurrence in open Metrosideros forest with mixed native understory. Preserved_Specimen Davis, J. 487 Hawaii BISH 1981-03-08
101 Geniostoma hedyosmifolium South Kona District; Honokau AHU Tr 65(40); W slopes of Mauna Loa; 1- 2 km north of Honokua 1950 lava flow Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Growing on pahoehoe. Preserved_Specimen Warshauer, F.R. 1924 Hawaii BISH 1978-07-01
102 Geniostoma hedyosmifolium Kawaipapa stream, above Nahiku-Hana area Metrosideros-Cheirodendron trigynum wet forest With Sadleria spp., Pipturus albidus, Clermontia grandiflora, Dubautia plantaginea, Cyrtandra grayana, Broussaisia arguta, Coprosma foliosa, Perrottetia grayana, Broussaisia arguta, Coprosma foliosa, Perrottetia sandwicensis, Clermontia kakeana, Polyscias oah Preserved_Specimen Perlman, S. 23127 Maui BISH 2012-11-14
103 Geniostoma hedyosmifolium below Halealoha Kipuka in 1855 flow Preserved_Specimen Forbes, C.N. 752 Hawaii BISH 1915-06-07
104 Geniostoma hedyosmifolium Napau Crater Rain forest. Preserved_Specimen Degener, O. 10245 Hawaii BISH 1929-12-05
105 Geniostoma hedyosmifolium South Hilo District; Waiakea AHU. Tr 28(121); east slope of Mauna Loa along power-line road; north of Stainback Highway Wet `ohi`a forest with other native trees, a treefern & native shrub understory. Preserved_Specimen Warshauer, F.R. 1274 Hawaii BISH 1977-08-02
106 Geniostoma hedyosmifolium Olinda, Pipe-line trail Dense, dark rain forest. Preserved_Specimen Degener, O. 10264 Maui BISH 1927-06-21
107 Geniostoma hedyosmifolium Kulani Prison Road (Stainback Hwy). 2 miles below prison and near water filled depression ca. 100 yards off the road Dense shade in tree fern-Metrosideros forest with Cyrtandra and extremely large Dumortiera. Preserved_Specimen Herbst, D.R. 368 Hawaii BISH 1967-03-26
108 Geniostoma hedyosmifolium South Hilo District; Makahanaloa AHU. Tr 20(46); east slope of Mauna Kea, along Kapue Stream; Hilo Forest Reserve Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 1471 Hawaii BISH 1977-09-05
109 Geniostoma hedyosmifolium Puna District; Olaa AHU. Tr 30(73); east slope of Mauna Loa; Upper Olaa Forest Reserve Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 1268 Hawaii BISH 1977-07-28
110 Geniostoma hedyosmifolium South Hilo District; Waiakea AHU. Tr 27(74); east slope of Mauna Loa at Tree Plant Road, 2.7 miles from stainback Highway Wet `ohi`a forest with other native trees, a treefern & native shrub understory. Preserved_Specimen Warshauer, F.R. 1720 Hawaii BISH 1978-03-03
111 Geniostoma hedyosmifolium Lahaina District, W. Maui Mtns. Along Eke Trail Diverse wet shrubland. Preserved_Specimen Ray, G. 1074 Maui BISH
112 Geniostoma hedyosmifolium East Maui, Waihoi Valley, Puu Hoolio Under Cheirodendron, in Dicranopteris & Paspalum conjugatum. Preserved_Specimen Harrison, B. 2520 Maui BISH 1972-07-20
113 Geniostoma hedyosmifolium East Maui, E Kopili ula, Acacia-Metrosideros, 50% open 7 m canopy, understory of uluhe mat ferns dissected by riparian veg with 'ohia & Cheirodendron, understory of Ilex, Clermontia arborescens, Scaevola chamissoniana, Myrsine lessertiana, Melicope spp, Hedyotis spp, Coprosma ssp, Perrottetia, and Broussaisia, shrubs of Vaccinium, Styphelia, Cyrtandra, and Rubus hawaiensis, ferns and bryophytes are common. Preserved_Specimen Wood, K.R. 11299 Maui BISH 2005-05-03
114 Geniostoma hedyosmifolium W Maui, Lahaina District, Haelaaua-Nakalalua Trail, 1 mile from Haelaaua Under Metrosideros trees. Preserved_Specimen Woolford, H.A. Maui BISH 1956-08-20
115 Geniostoma hedyosmifolium SW of Kaunuohua Closed Metrosideros forest with Cibotium, Pelea spp, Coprosma, Ilex, Gouldia, Broussaisia, Vaccinium and Cyrtandra. Preserved_Specimen Davis, J. 800 Molokai BISH 1982-05-23
116 Geniostoma hedyosmifolium Kahananui AHU Tr 10(4/5); on ridge S of Pakui Wet `ohi`a forest with other native trees and a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 2401 Molokai BISH 1979-07-11
117 Geniostoma hedyosmifolium Kilauea, Napau Trail, Chain of Craters In wet `ohi`a forest. Preserved_Specimen St.John, H. 11212 Hawaii BISH 1931-12-20
118 Geniostoma hedyosmifolium South Kona; Hoopuloa In dense forest Preserved_Specimen Rock, J.F.C. 25755 Hawaii BISH 1957-03-24
119 Geniostoma hedyosmifolium Waiakea; Site 27 OS Kulani Project Closed healthy forest. Soil a shallow, well drained drained organic over an a`a flow. Primary vegetation: Metrosideros polymorpha (pubescent), Ilex anomala, Myrsine lessertiana, Myrsine sandwicensis, Cheirodendron trigynum & Coprosma spp. Preserved_Specimen Adee, K.T. Hawaii BISH 1978-11-02
120 Geniostoma hedyosmifolium Kohala. Head of Alakahi Valley Rainswept shrubby plateau. Preserved_Specimen Degener, O. 19821 Hawaii BISH 1949-08-27
121 Geniostoma hedyosmifolium Waiakea; Upper Waiakea Forest Reserve, Site 12 OS Open healthy forest. Soil a shallow and well drained organic found over an a`a flow. Primary vegetation: Metrosideros polymorpha (sometimes pubescent), Ilex anomala, Myrsine lessertiana, M. sandwicensis & Cheirodendron trigynum. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-28
122 Geniostoma hedyosmifolium Wailulu District, Kahakuloa AHU Tr 38(7); West Maui mountains; in bog between Makamakaole & Huluhulupueo Strs Wet open bog with scattered native shrubs. Preserved_Specimen Warshauer, F.R. 3132 Maui BISH 1980-08-25
123 Geniostoma hedyosmifolium Kilauea, Chain of Craters between Makaopuhi and Napau Craters Wet forest Preserved_Specimen Fosberg, F.R. 10102 Hawaii BISH 1933-08-29
124 Geniostoma hedyosmifolium Along main road 17 miles from Kohala towards Waimea Ravine at elevation with considerable rainfall. Preserved_Specimen Degener, O. 30195 Hawaii BISH 1926-08-14
125 Geniostoma hedyosmifolium South Hilo District 4 miles northwest of Kulani Prison Camp Metrosideros forest. Preserved_Specimen St.John, H. 25324 Hawaii BISH 1954-12-19
126 Geniostoma hedyosmifolium Kawela AHU. Tr 5(1); Top of Papaala Pali, Kaunuohua Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Stemmermann, R.L. 3929 Molokai BISH 1979-06-21
127 Geniostoma hedyosmifolium Kahananui AHU. Tr 10(2); on northeast slope of ridge S of Pakui Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 2398 Molokai BISH 1979-07-11
128 Geniostoma hedyosmifolium E. Maui, Kipahulu valley, Palikea Stream, Delta camp region Acacia-Metrosideros-Cheirodendron montane wet forest with sections of Dicranopteris fernland. W/ Rubus hawaiiensis, Perrottetia, Broussaisia, Pipturus, Cyrtandra spathulata, Kadua affinis, Machaerina angustifolia, Psychotria, Diplazium sandwichianum, Clermontia arborescens, Freycinetia, Melico Preserved_Specimen Perlman, S. 13728 Maui BISH 2009-05-11
129 Geniostoma hedyosmifolium Puaahala AHU. Tr 9(5); Kalapamoa Ridge Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Stemmermann, R.L. 3872 Molokai BISH 1979-07-11
130 Geniostoma hedyosmifolium W Maui, Lahaina District, Ridge N of Amalu Stream. On margins Rain forest Preserved_Specimen Degener, O. 25059 Maui BISH 1959-03-25
131 Geniostoma hedyosmifolium W Maui, Puu Kukui Forest bog. Preserved_Specimen Munro, G.C. 610 Maui BISH 1916-09-25
132 Geniostoma hedyosmifolium Wailau, Ninole, Kaa Ahupuaa, SE slope of Mauna Loa, central Kau Forest between branches of Piikea Gulch. Transect 8, Station 64. Wet koa-ohia forest with other native trees and a treefern, native shrub understory. Preserved_Specimen Jacobi, J.D. 638 Hawaii BISH 1976-05-26
133 Geniostoma hedyosmifolium East Maui, Hana District, Waihoi Valley, S fork of Waiohonu Along stream bank with Vaccinium, ferns, mosses, etc. Preserved_Specimen Harrison, B. 78 Maui BISH 1972-07-13
134 Geniostoma hedyosmifolium E. Maui, Haleakala, Kipahulu Valley, SE Slope, 1/4 mi S. off Palikea stream On bank of small, intermittent stream Preserved_Specimen Medeiros, A.C. 450 Maui BISH 1983-07-09
135 Geniostoma hedyosmifolium Napau Crater Rain forest Preserved_Specimen Degener, O. 10243 Hawaii BISH 1930-01-25
136 Geniostoma hedyosmifolium Kohala Mountains summit On embankment Preserved_Specimen Rock, J.F.C. Hawaii BISH 1961-05-07
137 Geniostoma hedyosmifolium below Halealoha Kipuka in 1855 flow Preserved_Specimen Forbes, C.N. 752 Hawaii BISH 1915-06-07
138 Geniostoma hedyosmifolium Kona-Hilo Hwy, ca. 6.8 miles toward Hilo from Hongo Store Metrosideros-Cibotium forest community Preserved_Specimen Newell, C.L. 659 Hawaii BISH 1966-08-28
139 Geniostoma hedyosmifolium Piihonua; Hilo Forest Reserve, Site 05 OS Pubescent ohia dieback. Soil an extremely shallow & excessively drained organics over an a`a flow. Primary vegetation: Metrosideros polymorpha (pubescent), Ilex anomala, Myrsine lessertiana, M. sandwicensis & Cheirodendron trigynum. Preserved_Specimen Adee, K.T. Hawaii BISH
140 Geniostoma hedyosmifolium W Maui, Eke Crater On top in bog turf. Preserved_Specimen Medeiros, A.C. 328 Maui BISH 1982-11-09
141 Geniostoma hedyosmifolium S slope of Mauna Loa, above Pahala Forest. Preserved_Specimen Skottsberg, C.J.F. 590 Hawaii BISH 1922-09-19
142 Geniostoma hedyosmifolium East Maui, Makawao District, Kalialinui AHU, Tr 2(8), NW slope of Haleakala, on upper ridge between Waiohiwi & Kahakapo Mesic koa-`ohi`a forest with other native trees, a native fern and shrub understorey. Preserved_Specimen Warshauer, F.R. 2724 Maui BISH 1980-06-28
143 Geniostoma hedyosmifolium Kilauea Iki Trail In `ohi`a forest near trail Preserved_Specimen Fagerlund, G.O. 143 Hawaii BISH 1942-11-08
144 Geniostoma hedyosmifolium Kehena Ditch Trail Near bottom of first gulch (running stream) west of Honokanenui Metrosideros-Cheirodendron forest, with Pelea, Vaccinium, patches of Spaghnum, Dubautia, Gouldia, Straussia, Clermontia, Gunnera & Cyanea. Preserved_Specimen Gillett, G.W. 1904 Hawaii BISH 1966-06-26
145 Geniostoma hedyosmifolium Lahaina DISTR. Honokowai AHU. Tr 36(22); Along Pu'ukukui Trail, W Maui Forest Reserve Wet ohia forest with other native trees and a treefern and native shrub understory. Preserved_Specimen Higashino, P.K. 9486 Maui BISH 1980-06-16
146 Geniostoma hedyosmifolium Along Saddle Road 16 miles from Hilo In rich, forested kipuka south of road. Preserved_Specimen Degener, O. 19793 Hawaii BISH 1949-08-30
147 Geniostoma hedyosmifolium Puna District; Olaa AHU. Tr 29(91); east slope of Mauna Loa, S of Puu Makaala; Olaa Forest Reserve Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 1658 Hawaii BISH 1978-01-06
148 Geniostoma hedyosmifolium East Maui, Waihoi Valley, Puu Hoolio Under Cheirodendron, in Dicranopteris & Paspalum conjugatum. Preserved_Specimen Harrison, B. 2520 Maui BISH 1972-07-20
149 Geniostoma hedyosmifolium Papaikou, E slope of Muana Kea, just N of Kapue Stream. Transect 20, Station 50. Wet ohia forest with other native trees and a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 1495 Hawaii BISH 1977-10-15
150 Geniostoma hedyosmifolium East Maui, E Kopili ula Acacia-Metrosideros, 50% open 7 m canopy, understory of uluhe mat ferns dissected by riparian veg with 'ohia & Cheirodendron, understory of Ilex, Clermontia arborescens, Scaevola chamissoniana, Myrsine lessertiana, Melicope spp, Hedyotis spp, Corposma spp, Perrottetia, and Broussaisia, shrubs of Vaccinium, Styphelia, Cyrtandra, and Rubus hawaiensis, ferns and bryophytes are common. Preserved_Specimen Wood, K.R. 11288 Maui BISH 2005-05-03
151 Geniostoma hedyosmifolium Lahaina District, Honolua AHU, Tr 35(16); Ridge between Honolua & Honokohau Valleys; West Maui Forest Reserve Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Higashino, P.K. 9478 Maui BISH 1980-08-15
152 Geniostoma hedyosmifolium Puna District. Fern Forest Vacation Estates Rain forest Preserved_Specimen Gagné, W.C. 600 Hawaii BISH 1971-04-04
153 Geniostoma hedyosmifolium Wailau AHU. Tr 19(24); Between Wailau & Pelekunu Valleys; along Kapapa Pali Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Higashino, P.K. 9433 Molokai BISH 1980-08-03
154 Geniostoma hedyosmifolium Up Kulani Prison Road, 5.3 miles toward Prison from intersection with Volcano Road Metrosideros-Cibotium forest community. Preserved_Specimen Newell, C.L. 694 Hawaii BISH 1966-09-01
155 Geniostoma hedyosmifolium South Kohala District; Awini AHU. Tr 78(10/9); Kohala Mountains; southwest of Honokea & northeast of Honokanenui Streams Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Preserved_Specimen Warshauer, F.R. 2483 Hawaii BISH 1979-08-01
156 Geniostoma hedyosmifolium Piihonua; Hilo Forest Reserve, on the trail to Site 02 OS Stunted ohia wetland dieback; soil a shallow poorly drained organic type found on a pahoehoe flow. One small area of water was at or near the surface. Primary vegetation: Metrosideros polymorpha, Ilex anomala, Gouldia sp. & Pelea clusiifolia. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-05
157 Geniostoma hedyosmifolium Puu Huluhulu Deep samll crater within Pu`u. Growing inside crater with Cibotium, Stenogyne rugosa, Cyrtandra platyphylla, Clermontia hawaiiensis, Peperomia spp., and Metrosideros. Preserved_Specimen Davis, J. 451 Hawaii BISH 1981-02-15
158 Geniostoma hedyosmifolium Waiakea; Site 20 OS Upper Waiakea Forest Reserve Closed healthy forest. Shallow well drained organic soil over pahoehoe flow. Primary vegetation: Metrosideros polymorpha, Ilex anomala, Myrsine lessertiana, M. sandwicensis, Cheirodendron trigynum & Psychotria hawaiiensis. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-05-29
159 Geniostoma hedyosmifolium Keauhou Ranch, Kau District, near jeep trail connecting Kilauea Forest Reserve boundary and powerline trail Tall closed Metrosideros forest in kipuka surrounded by more recent lava. Prostrate on ground under o`hia. Preserved_Specimen Cuddihy, L.W. 48 Hawaii BISH 1979-12-12
160 Geniostoma hedyosmifolium Kona-Hilo Hwy, ca. 6.8 miles toward Hilo from Hongo Store Metrosideros-Cibotium forest community Preserved_Specimen Newell, C.L. 655 Hawaii BISH 1966-08-28
161 Geniostoma hedyosmifolium W Maui, Lahaina District, Haelaau, above Honokowai Wooded ridge Preserved_Specimen St.John, H. 10228 Maui BISH 1930-02-06
162 Geniostoma hedyosmifolium Hana District. Heleleokaoha stream area, Huki camp of fence crew, above Hana on East Maui . Metrosideros-Dicranopteris linearis wet forest With Sadleria spp., Pipturus albidus, Clermontia grandiflora, Dubautia plantaginea, Cyrtandra grayana, Broussaisia arguta, Coprosma foliosa, Perrottetia sandwicensis, Clermontia kakeana, Polyscias oah Preserved_Specimen Perlman, S. 23968 Maui BISH 2014-05-28
163 Geniostoma hedyosmifolium East Maui, Hana District, Waihoi Valley, S fork of Waiohonu, Ridge above stream In uluhe (fern), under Cheirodendron & Metrosideros Preserved_Specimen Harrison, B. 8 Maui BISH 1972-06-14
164 Geniostoma hedyosmifolium Kau Forest Reserve, Kau District Open Metrosideros - Acacia koa forest with closed Cibotium understory. Preserved_Specimen Davis, J. 648 Hawaii BISH 1981-11-19
165 Geniostoma hedyosmifolium Olaa; Site 24 OS Olaa Forest Reserve Closed healthy forest. Soil a moderately deep & well drained organic/ash over a pahoehoe flow. Primary vegetation: Metrosideros polymorpha, Ilex anomala, Myrsine lessertiana, Cheirodendron trigynum, Coprosma spp. & Psychotria hawaiiensis. Preserved_Specimen Adee, K.T. Hawaii BISH 1978-09-28
166 Geniostoma hedyosmifolium Olaa; Site 31 OS Olaa Forest Reserve Closed healthy forest. Soil a moderately deep & well drained organic/ash over a pahoehoe flow. Primary vegetation: Metrosideros polymorpha, Ilex anomala, Myrsine lessertiana, Cheirodendron trigynum, Coprosma spp. & Psychotria hawaiiensis. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-01-30
167 Geniostoma hedyosmifolium W Maui, Mountain of W Maui Preserved_Specimen Mann, H. 434 Maui BISH 1864-01-01
168 Geniostoma hedyosmifolium Puna District; Olaa AHU. Tr 28(145); east slope of Mauna Loa; S of Stainback highway; Olaa Forest Reserve Wet exotic tree, `ohi`a forest with a matted-fern & native shrub understory. Preserved_Specimen Warshauer, F.R. 1280 Hawaii BISH 1977-08-04
169 Geniostoma hedyosmifolium South Hilo District; Waiakea AHU. Tr 28(88); east slope of Mauna Loa across jeep road north of Stainback Hwy Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Growing on pahoehoe. Preserved_Specimen Warshauer, F.R. 1644 Hawaii BISH 1977-12-31
170 Geniostoma hedyosmifolium East Maui, Hana District, Tr 11(31), NE slope of Haleakala, Hana FR; leaving ridge going toward str Wet `ohi`a forest with other native trees, a treefern and native shrub understory. Growing on `a`a substrate. Preserved_Specimen Higashino, P.K. 9324 Maui BISH 1980-06-30
171 Geniostoma hedyosmifolium W Maui, Lahaina District, Puu Kukui, between Haelaau & Nakalalua Moist forest. Preserved_Specimen Fosberg, F.R. 10044 Maui BISH 1933-08-24
172 Geniostoma hedyosmifolium Ridge rt side Kipahulu Preserved_Specimen Forbes, C.N. 1688 Maui BISH 1919-11-20
173 Geniostoma hedyosmifolium East Maui, Honomanu Preserved_Specimen Rock, J.F.C. 8538 Maui BISH 1910-09-01
174 Geniostoma hedyosmifolium W Maui, Honokawai Preserved_Specimen Rock, J.F.C. 8185 Maui BISH 1910-08-01
175 Geniostoma hedyosmifolium W Maui, Puu Kukui Preserved_Specimen Munro, G.C. 620 Maui BISH 1916-09-25
176 Geniostoma hedyosmifolium W Maui, Honokahua-Amalu ridge above Haelaau Preserved_Specimen Cranwell, L.M. 3681 Maui BISH 1938-08-01
177 Geniostoma hedyosmifolium East Maui, Honomanu Preserved_Specimen Rock, J.F.C. 10344 Maui BISH 1911-05-01
178 Geniostoma hedyosmifolium Nalehu forests Preserved_Specimen Rock, J.F.C. 10015 Hawaii BISH 1912-01-01
179 Geniostoma hedyosmifolium Wailuku Pali Preserved_Specimen Forbes, C.N. 2443 Maui BISH 1920-05-21
180 Geniostoma hedyosmifolium Disappointment Trail, off of Stainback Hwy. In forest at beginning of trail Preserved_Specimen Wagner, W.L. 5594 Hawaii BISH 1985-06-26
181 Geniostoma hedyosmifolium East Maui, Lower Shelf of Kipahulu Valley above Delta Camp in snare group BL-60 Preserved_Specimen Welton, P. 170 Maui BISH 1993-06-30
182 Geniostoma hedyosmifolium Honokauenui, Kohala Preserved_Specimen Rock, J.F.C. 8364 Hawaii BISH 1910-06-01
183 Geniostoma hedyosmifolium W Maui, Eke Crater Preserved_Specimen Medeiros, A.C. 333 Maui BISH 1982-11-09
184 Geniostoma hedyosmifolium W Maui, Puu Kukui, Below summit bog Preserved_Specimen Cranwell, L.M. 3681 Maui BISH 1938-07-26
185 Geniostoma hedyosmifolium Main ridge of Lanai Preserved_Specimen Rock, J.F.C. 8017 Lanai BISH 1910-07-25
186 Geniostoma hedyosmifolium Waimea Alakahi-Kawainui Preserved_Specimen Rock, J.F.C. 4301 Hawaii BISH 1909-07-12
187 Geniostoma hedyosmifolium Lanaihale Preserved_Specimen Rock, J.F.C. 8041 Lanai BISH 1910-07-25
188 Geniostoma hedyosmifolium Kipahulu Valley Ridge, left side Preserved_Specimen Forbes, C.N. 1644 Maui BISH 1910-05-01
189 Geniostoma hedyosmifolium Waimea Preserved_Specimen Rock, J.F.C. 8316 Hawaii BISH 1910-06-01
190 Geniostoma hedyosmifolium W Maui, Mountain of W Maui Preserved_Specimen Mann, H. 434 Maui BISH 1864-01-01
191 Geniostoma hedyosmifolium Alakahi-Kawainui Gorge Preserved_Specimen Rock, J.F.C. 4301 Hawaii BISH 1909-07-12
192 Geniostoma hedyosmifolium Pepeopae Bog to edge of Pelekunu Valley Preserved_Specimen Herbst, D.R. 2958 Molokai BISH 1973-04-04
193 Geniostoma hedyosmifolium W Maui, Maunahooma (?) Preserved_Specimen Forbes, C.N. 9 Maui BISH 1910-05-01
194 Geniostoma hedyosmifolium Kona; Kealapuali Preserved_Specimen Forbes, C.N. 298 Hawaii BISH 1911-06-30
195 Geniostoma hedyosmifolium East Maui, Nahiku Preserved_Specimen Forbes, C.N. 257 Maui BISH 1920-07-01
196 Geniostoma hedyosmifolium W Maui, Puu Kukui Preserved_Specimen Gillett, G.W. 2426 Maui BISH 1972-08-02
197 Geniostoma hedyosmifolium East Maui, Makawao Forest Reserve Preserved_Specimen Woolford, H.A. 179 Maui BISH 1956-01-01
198 Geniostoma hedyosmifolium Olinda Pipe Line Preserved_Specimen Woolford, H.A. 177 Maui BISH 1956-01-01
199 Geniostoma hedyosmifolium E. Maui, Kipahulu Valley, Lower Valley Preserved_Specimen Medeiros, A.C. 443 Maui BISH 1983-07-09
200 Geniostoma hedyosmifolium Lanaihale, highest ridge on Lanai Preserved_Specimen Rock, J.F.C. 8020 Lanai BISH 1910-07-22
201 Geniostoma hedyosmifolium Kohala Preserved_Specimen Rock, J.F.C. 8430 Hawaii BISH 1910-06-01
202 Geniostoma hedyosmifolium Hualalai, West slope, Honokahau. Frank Greenwell's paddocks Preserved_Specimen Kondo, Y. 0.333 Hawaii BISH 1946-07-26
203 Geniostoma hedyosmifolium Kipahulu Preserved_Specimen Forbes, C.N. 1688 Maui BISH 1919-11-20
204 Geniostoma hedyosmifolium East Maui, Honomanu Preserved_Specimen Rock, J.F.C. 10344 Maui BISH 1911-05-01
205 Geniostoma hedyosmifolium W Maui, Honokawai Preserved_Specimen Rock, J.F.C. 8155 Maui BISH 1910-08-01
206 Geniostoma hedyosmifolium W Maui, Honokawai Preserved_Specimen Rock, J.F.C. 8153 Maui BISH 1910-08-01
207 Geniostoma hedyosmifolium West side of Haipuaena Preserved_Specimen Forbes, C.N. 2650 Maui BISH 1920-06-28
208 Geniostoma hedyosmifolium East Maui, Hana District, Kipahulu Valley, W part Preserved_Specimen Lamoureux, C.H. 3994 Maui BISH 1967-08-07
209 Geniostoma hedyosmifolium Waimea, Alakahi-Kawainui Ditch Trail Preserved_Specimen Rock, J.F.C. 4305 Hawaii BISH 1909-07-12
210 Geniostoma hedyosmifolium East bank of Honomanu Gulch Preserved_Specimen Forbes, C.N. 2565 Maui BISH 1920-06-21
211 Geniostoma hedyosmifolium E Maui, Kipahulu Valley, Upper Valley Preserved_Specimen Medeiros, A.C. 455 Maui BISH 1983-07-10
212 Geniostoma hedyosmifolium Waimea, Alakahi-Kawainui Ditch Trail Preserved_Specimen Rock, J.F.C. 4305 Hawaii BISH 1909-07-12
213 Geniostoma hedyosmifolium West Maui, Lihau Preserved_Specimen Hobdy, R.W. 3302 Maui BISH 1991-06-04
214 Geniostoma hedyosmifolium Between Halealoha & Halelouolu Preserved_Specimen Forbes, C.N. 726 Hawaii BISH 1915-06-06
215 Geniostoma hedyosmifolium Lanaihale, highest ridge of Lanai Preserved_Specimen Rock, J.F.C. 8020 Lanai BISH 1910-07-22
216 Geniostoma hedyosmifolium Trail between Wailuku River & Halelouolu Preserved_Specimen Forbes, C.N. 721 Hawaii BISH 1915-06-05
217 Geniostoma hedyosmifolium Nalehu forests Preserved_Specimen Rock, J.F.C. 10015 Hawaii BISH 1912-01-01
218 Geniostoma hedyosmifolium W Maui, Puu Kukui Preserved_Specimen Woolford, H.A. Maui BISH 1957-08-29
219 Geniostoma hedyosmifolium E Maui, Kipahulu Valley, Upper Valley Preserved_Specimen Medeiros, A.C. 456 Maui BISH 1983-07-10
220 Geniostoma hedyosmifolium Kohala Preserved_Specimen Rock, J.F.C. 8430 Hawaii BISH 1910-06-01
221 Geniostoma hedyosmifolium East Maui, Waihoi Bog On margins Preserved_Specimen Hobdy, R.W. 2582 Maui BISH 1986-07-11
222 Geniostoma hedyosmifolium W Maui, Iao Valley Preserved_Specimen Forbes, C.N. 102 Maui BISH 1910-06-01
223 Geniostoma hedyosmifolium Kipahulu Valley, Haleakala National Park, National Park Service Expedition III, West camp, transect 4 Preserved_Specimen Russell, C.A. 543 Maui BISH 1984-03-02
224 Geniostoma hedyosmifolium Lanaihale, highest ridge of Lanai Preserved_Specimen Rock, J.F.C. 8020 Lanai BISH 1910-07-22
225 Geniostoma hedyosmifolium Kilohana In flow of 1855 Preserved_Specimen Forbes, C.N. 723 Hawaii BISH 1915-06-06
226 Geniostoma hedyosmifolium Manawainui Gulch. At left fork Preserved_Specimen Herbst, D.R. 3026 Maui BISH 1973-05-24
227 Geniostoma hedyosmifolium W Maui, Honokawai Preserved_Specimen Rock, J.F.C. 8153 Maui BISH 1910-08-01
228 Geniostoma hedyosmifolium East Maui, Honomanu Preserved_Specimen Rock, J.F.C. 10344 Maui BISH 1911-05-01
229 Geniostoma hedyosmifolium Hawaii or Maui Preserved_Specimen Hillebrand, W.B. BISH
230 Geniostoma hedyosmifolium Upper Waiakea Forest Reserve. Site 357-26 Preserved_Specimen Adee, K.T. Hawaii BISH
231 Geniostoma hedyosmifolium Honaunau forest Preserved_Specimen Sakai, H. Hawaii BISH 1980-06-18
232 Geniostoma hedyosmifolium Honaunau Forest Reserve Preserved_Specimen Bryan, L.W. Hawaii BISH 1957-03-01
233 Geniostoma hedyosmifolium Honolii Preserved_Specimen Adee, K.T. Hawaii BISH 1978-03-07
234 Geniostoma hedyosmifolium South Hilo District, Waiakea. Buffer zone collections along Stainback Highway Preserved_Specimen Funk, E.J. Hawaii BISH 1998-12-15
235 Geniostoma hedyosmifolium Hawaii Preserved_Specimen Rémy, J. Hawaii BISH 1851-01-01
236 Geniostoma hedyosmifolium West Maui, Eeke-Honokahau Preserved_Specimen Rock, J.F.C. Maui BISH 1918-08-01
237 Geniostoma hedyosmifolium Waihee Preserved_Specimen Lydgate, J.M. Maui BISH 1870-08-01
238 Geniostoma hedyosmifolium Lower Kipahulu Valley, Area between upper pasture and the gauging station on Palakea Stream, for Resources Basic Inventory Preserved_Specimen Gagné, B.H. Maui BISH 1975-03-01
239 Geniostoma hedyosmifolium East Maui, Nahiku Stream Preserved_Specimen Rock, J.F.C. Maui BISH 1911-05-01
240 Geniostoma hedyosmifolium East Maui, Honomanu Preserved_Specimen Rock, J.F.C. Maui BISH 1911-05-01
241 Geniostoma hedyosmifolium Kona. Honaunau forest Preserved_Specimen Carlson, N.K. Hawaii BISH 1967-06-01
242 Geniostoma hedyosmifolium Hawaii Preserved_Specimen Rock, J.F.C. Hawaii BISH
243 Geniostoma hedyosmifolium Lanai Preserved_Specimen Hillebrand, W.B. Lanai BISH
244 Geniostoma hedyosmifolium Kipahulu Valley, along Palikea Stream. Above the waterfall Preserved_Specimen Warner, R.E. 17 Maui BISH 1967-08-01
245 Geniostoma hedyosmifolium Manawainui Gulch. At left fork Preserved_Specimen Herbst, D.R. 3025 Maui BISH 1973-05-24
246 Geniostoma hedyosmifolium Waimea Preserved_Specimen Rock, J.F.C. 8316 Hawaii BISH 1910-06-01
247 Geniostoma hedyosmifolium Eke; Honokahau Drainage Basin Preserved_Specimen Forbes, C.N. 387 Maui BISH 1917-09-25
248 Geniostoma hedyosmifolium Lanaihale, highest ridge on Lanai Preserved_Specimen Rock, J.F.C. 8020 Lanai BISH 1910-07-22
249 Geniostoma hedyosmifolium W Maui, Hanakaoo Preserved_Specimen Forbes, C.N. 42 Maui BISH 1910-05-01
250 Geniostoma hedyosmifolium Mountains above Puu Kolekole Preserved_Specimen Forbes, C.N. 204 Molokai BISH 1912-06-01
251 Geniostoma hedyosmifolium Waimea, Kohala Mountains Preserved_Specimen Forbes, C.N. 509 Hawaii BISH 1911-09-01
252 Geniostoma hedyosmifolium East maui, Lower shelf of kipahulu valley above delta in snare group BH-157 Preserved_Specimen Welton, P. 1878002 Maui BISH 1995-04-22
253 Geniostoma hedyosmifolium Hawaii Preserved_Specimen Rémy, J. 362 Hawaii BISH 1851-01-01
254 Geniostoma hedyosmifolium Olaa Forest Reserve, behind Volcano dump Preserved_Specimen Wagner, W.L. 5721 Hawaii BISH 1984-02-28
255 Geniostoma hedyosmifolium Puna District, Hawaiina Orchid Isle Estates Open canopy Metrosideros forest. Preserved_Specimen Davis, J. 488 Hawaii BISH 1981-03-08
256 Geniostoma hedyosmifolium Kilauea, Chain of Craters between Makaopuhi and Napau Craters Wet forest Preserved_Specimen Fosberg, F.R. 10102 Hawaii BISH 1933-08-29
257 Geniostoma hedyosmifolium Piihonua; Hilo Forest Reserve, Site 13 OS Pubescent ohia dieback. Soil an extremely shallow and excessively drained organic type found over an a'a flow. Primary vegetation: Metrosideros polymorpha (pubescent), Ilex anomala, Myrsine lessertiana, M. sandwicensis & Cheirodendron trigynum. Preserved_Specimen Adee, K.T. Hawaii BISH 1979-06-26