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Nephrolepis exaltata

(L.) Schott (1834)

This name is accepted

Kingdom: Viridiplantae Phylum: Pteridophyta Class/Clade: Polypodiopsida Order: Polypodiales Family: Nephrolepidaceae Genus: Nephrolepis

‘ōkupukupu [okupukupu], ‘ōkupukupu [okupukupu], kupukupu, ni‘ani‘au [nianiau], ni‘ani‘au [nianiau], palapalai, pāmoho [pamoho], sword fern

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Description

Key Characters:

Growth Form:

Stems:

Roots:

Leaves:

Sori:

Gametophyte:

Ploidy:

Habitat:

Elevation Range:

Historical Distribution

Images

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Uses and Culture

USES

  • Sometimes used on hula altars (kuahu) (noted for Maui in Pukui 1942).

PROPAGATION/CULTIVATION

  • Easy from small plant. Few pests. (Bornhorst 1996:73–74). Best under medium to high light in moist to dry soils (Hoshizaki & Moran 2002:393).

Natural History

Statewide Status

Indigenous

Island Status

Ni'ihau(Incl. Lehua) Indigenous
Kaua'i Indigenous
O'ahu Indigenous
Molokai Indigenous
Lana'i Indigenous
Kaho'olawe Potentially Naturalizing
Maui Indigenous
Hawai'i Indigenous

Dispersal Agents


Bibliography

Name Published In: Gen. Fil.: t. 3 (1834)

Other References

Palmer 2003:190 (Ni, K, O, Mo, L, M, H [as N. e. subsp. hawaiiensis]); Staples & Herbst 2005:40 (KEY), 41 (DESCR); Hovenkamp & Miyamoto 2005:300/Palmer & Thomas in press (Syn. N. e. subsp. hawaiiensis = N. exaltata, status change from end to ind); Ranker et al. 2019:68 (maintain endemic subsp. hawaiiensis, not followed); Paul Higashino pers. comm. 2025 (Ka? [observed but no spm. under rain catchment]); Herbst 4037 HAW (L†, 1974)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Nephrolepis exaltata Waimanalo Shady, wet mesic forest. Growing right along trail. Preserved_Specimen Arthur, K. 25 Oahu BISH 2025-01-23
2 Nephrolepis exaltata Kamakou Preserve, gulch W of Puu o Kaeha Fraxinus ohdei, Osmanthus, Dodonoea, Alyxia, Dubautia Preserved_Specimen Higashino, P.K. 10089 Molokai BISH 1983-10-11
3 Nephrolepis exaltata W ridge of Honomuni dry thicket Preserved_Specimen St.John, H. 25215 Molokai BISH 1953-12-23
4 Nephrolepis exaltata Hauula dry sunny embankment Preserved_Specimen Degener, O. 3220 Oahu BISH 1922-08-16
5 Nephrolepis exaltata Kilauea, Kipuka Puaulu open woodland, lush grass, deep soil Preserved_Specimen Fosberg, F.R. 33272 Hawaii BISH 1951-02-11
6 Nephrolepis exaltata Puna, near RR track flow of 1840 Preserved_Specimen Forbes, C.N. 1059 Hawaii BISH 1915-07-07
7 Nephrolepis exaltata Kalalau Valley, N side, E of Keanapuka Falls matting fern ridge, along drainage; Dicranopteris, Diplopterygium, Santalum freycinetianum, Kadua affinis, Coprosma waimeae, Metrosideros polymorpha, Dodonaea viscosa, Dianella sandwicensis, Pteridium decompositum, Scaevola procera, Xylosma hawaiiense, Psychotria mariniana, Ilex anomala, Bobea elatior, Pouteria sandwicensis, Syzygium sandwicensis Preserved_Specimen Wood, K.R. 12474 Kauai BISH 2007-07-30
8 Nephrolepis exaltata West ridge of Honomuni. Moist metrosideros forest. Preserved_Specimen St.John, H. 25212 Molokai BISH 1953-12-23
9 Nephrolepis exaltata Puna, lower Pulama in lava crack, shade, damp Preserved_Specimen Stone, B.C. 3008 Hawaii BISH 1959-07-02
10 Nephrolepis exaltata Lyon Arboretum, nose of ridge above section 10 on koa Preserved_Specimen Baker, R.F. 128 Oahu BISH 1975-06-13
11 Nephrolepis exaltata Palolo Valley koa forest Preserved_Specimen Fosberg, F.R. 9279 Oahu BISH 1933-03-19
12 Nephrolepis exaltata E hairpin turn Pali Road Mango grove Preserved_Specimen Topping, D.L. 2668 Oahu BISH 1923-06-17
13 Nephrolepis exaltata Olokui, Wailau, ridge between upper forks of Wailele Stream moist woods Preserved_Specimen St.John, H. 23325 Molokai BISH 1948-03-06
14 Nephrolepis exaltata Puna, Kaueleau Pandanus grove Preserved_Specimen St.John, H. 22285 Hawaii BISH 1946-12-22
15 Nephrolepis exaltata Waimea District. Kumuwela ridge, along road. Metrosideros - Acacia mesic forest Leptecophylla tameamea, Coprosma waimeae, Dodonaea viscosa, Wikstroemia furcata, Melicope ovata, M. barbigera, Cyanea leptostegia, Claoxylon sandwicense, Planchonella sandwicensis Preserved_Specimen Wood, K.R. 15453 Kauai BISH 2013-03-29
16 Nephrolepis exaltata Haupu, Kipu edge of forest Preserved_Specimen St.John, H. 13641 Kauai BISH 1933-12-25
17 Nephrolepis exaltata Kilauea fern jungle Preserved_Specimen Skottsberg, C.J.F. 3325 Hawaii BISH 1938-09-17
18 Nephrolepis exaltata Chain of Craters Rd. ~9 mi from intersection with Crater Rim Rd.; up Old Kalapana Trail ~0.3 mi.; ~10 m to L of trail Metrosideros-Dicranopteris Preserved_Specimen Newell, C.L. 560 Hawaii BISH 1966-08-23
19 Nephrolepis exaltata SW flank of Mauna Loa, Manuka open Metrosideros forest on aa with Nephrolepis, Sadleria, Myrsine, Styphelia; aa rocks covered with Cladina, Cladonia, Racomitrium Preserved_Specimen Higashino, P.K. 5933 Hawaii BISH 1977-07-03
20 Nephrolepis exaltata W Maui, Lahaina District, Lihau Montane Ohia Mixed Wet Forest at 4185 ft under a Metrosideros canopy with Coprosma, Hedyotis terminalis, Cibotium, Myrsine, Vaccinium, Psychotria, Ilex, Sadleria souleyetiana and a ground cover of Liparis, Peperomia, native ferns, moss and seedlings Preserved_Specimen Welton, P. 1187 Maui BISH 1991-08-25
21 Nephrolepis exaltata S. Hilo, ~17 km on Saddle Rd along thick growth of Dicranopteris Preserved_Specimen Herat, T. 697 Hawaii BISH 1974-06-22
22 Nephrolepis exaltata Olomana Moist Aleurites forest. Preserved_Specimen Christophersen, E. Oahu BISH 1924-10-26
23 Nephrolepis exaltata 2.5 mi W of Waihee, coast W of Wailuku steep banks above road Preserved_Specimen Fosberg, F.R. 56717 Maui BISH 1977-03-18
24 Nephrolepis exaltata Waianae Mts., Waianae Kai Trail, left side of trail on way up , above junction Alien mixed mesic forest with Kukui (Aleurites moluccana), Guava (Psidium cattleianum), and Silk oak (Grevillea robusta). Preserved_Specimen Crago, L.M. 2004001 Oahu BISH 2004-11-09
25 Nephrolepis exaltata Koolau Mts., Kona, Honolulu FR, Manoa Cliffs Trail, near Hawaiian Telephone Co. facility along paved road Preserved_Specimen Lammers, T.G. 5311 Oahu BISH 1983-08-27
26 Nephrolepis exaltata Kipapa Gulch, Waipio Metrosideros forest Preserved_Specimen Grant, M.L. 7178 Oahu BISH 1934-08-06
27 Nephrolepis exaltata Molokai Community College Farm shadehouse, in 1-gallon pot. Preserved_Specimen Evans, M. 60 Molokai BISH 1990-07-06
28 Nephrolepis exaltata Mapulehu Valley walls of rocky gorge Preserved_Specimen St.John, H. 12771 Molokai BISH 1932-12-28
29 Nephrolepis exaltata Koolau Mts., Laie Ridge trail partly shady Preserved_Specimen Herat, T. 366 Oahu BISH 1973-06-28
30 Nephrolepis exaltata Wainiha Valley dry, along trail Preserved_Specimen St.John, H. 13903 Kauai BISH 1934-01-01
31 Nephrolepis exaltata Kaneohe, Oahu Marugame Nursery, Continually damp greenhouse. Preserved_Specimen Lau, J. 1111 Oahu BISH 1985-06-17
32 Nephrolepis exaltata Puu Kanehoa, Honouliuli wooded ridge Preserved_Specimen St.John, H. 11061 Oahu BISH 1931-03-22
33 Nephrolepis exaltata E of Puu Kamana, S ridge of Kipapa Gulch, Waipio, Koolau Mts moist Preserved_Specimen St.John, H. 11637 Oahu BISH 1932-05-15
34 Nephrolepis exaltata below Palehua, Waianae Mts wooded valley, on ground Preserved_Specimen Hume, E.P. 322 Oahu BISH 1931-11-22
35 Nephrolepis exaltata Maunapuluo, Hanakapiai, Napali Coast on cliff Preserved_Specimen St.John, H. 10886 Kauai BISH 1930-12-01
36 Nephrolepis exaltata Chain of Craters Rd. ~9 mi from intersection with Crater Rim Rd.; up Old Kalapana Trail ~0.3 mi.; ~10 m to L of trail Metrosideros-Dicranopteris Preserved_Specimen Newell, C.L. 561 Hawaii BISH 1966-08-23
37 Nephrolepis exaltata gully near Haleakala Sapindus oahuensis, Rauvolfia sandwicensis, Antidesma pulvinatum; dry stream bed Preserved_Specimen Takeuchi, W.N. 3310 Oahu BISH 1987-01-01
38 Nephrolepis exaltata Paunau-Kuhua divide moist woods Preserved_Specimen St.John, H. 17698 Maui BISH 1936-12-19
39 Nephrolepis exaltata end of Hilina Pali Rd.; down trail ~1.5 mi open Metrosideros, Diospyros forest on aa lava Preserved_Specimen Newell, C.L. 436 Hawaii BISH 1966-08-18
40 Nephrolepis exaltata Puu Kanehoa, Waianae Mts., Honouliuli wet forest Preserved_Specimen Inafuku, R. Oahu BISH 1931-03-22
41 Nephrolepis exaltata Mt. Tantalus upper woods Preserved_Specimen St.John, H. 9879 Oahu BISH 1929-09-26
42 Nephrolepis exaltata Oahu on tree trunks Preserved_Specimen Judd, J.R. Oahu BISH 1888-01-01
43 Nephrolepis exaltata Kohala Mts., Awini Ridge, PTBG preserve on tussocks or on ohia Preserved_Specimen Flynn, T.W. 1376 Hawaii BISH 1985-11-19
44 Nephrolepis exaltata Koolau Mts., windward side, just below Nuuanu Pali on rocks slightly covered with soil Preserved_Specimen Harris, J.A. 242021 Oahu BISH 1924-08-14
45 Nephrolepis exaltata E side of Kamilonui valley, Maunalua rock ledge Preserved_Specimen Fosberg, F.R. 13605 Oahu BISH 1937-03-07
46 Nephrolepis exaltata Palolo, Kaau crater on ohia in bog Preserved_Specimen Chock, A.K. 419 Oahu BISH 1952-02-12
47 Nephrolepis exaltata East Maui, Waihoi Valley, N side with other ferns on logs Preserved_Specimen Harrison, B. 2594 Maui BISH 1972-11-21
48 Nephrolepis exaltata Koolau Mts., Kaluanui Stream Preserved_Specimen Newhouse, W.J. 53151 Oahu BISH 1953-10-18
49 Nephrolepis exaltata Mt. Kalena, Waianae Preserved_Specimen Grant, M.L. 7389 Oahu BISH 1934-09-30
50 Nephrolepis exaltata Nuuanu Valley Preserved_Specimen Mann, H. 547 Oahu BISH 1864-01-01
51 Nephrolepis exaltata S Kona, Kapua, Kaulanamauna, Manuka Preserved_Specimen Higashino, P.K. 6558 Hawaii BISH 1977-06-01
52 Nephrolepis exaltata Palolo Valley Preserved_Specimen Garber, D.W. 36 Oahu BISH 1919-11-02
53 Nephrolepis exaltata Pauoa Valley Preserved_Specimen Mann, H. 142393 Oahu BISH 1864-01-01
54 Nephrolepis exaltata Upper Palolo; south rim of Kaau Crater Preserved_Specimen Long, C.R. 3687 Oahu BISH 1965-12-30
55 Nephrolepis exaltata peak of Mt. Olympus Preserved_Specimen Garber, D.W. 293 Oahu BISH 1920-03-07
56 Nephrolepis exaltata Keawapilau; Behind releve 11 Preserved_Specimen Welton, P. 738 Oahu BISH 1991-06-27
57 Nephrolepis exaltata East Maui, Koolau Gap, E side Preserved_Specimen Hobdy, R.W. 3383 Maui BISH 1991-07-18
58 Nephrolepis exaltata West Maui, Lihue Preserved_Specimen Hobdy, R.W. 3299 Maui BISH 1991-06-04
59 Nephrolepis exaltata Nuuanu Valley Preserved_Specimen Mann, H. 547 Oahu BISH 1864-01-01
60 Nephrolepis exaltata East Maui, Hana District, SW side of Haleakala, Lualailua Preserved_Specimen Forbes, C.N. 2108 Maui BISH 1920-03-25
61 Nephrolepis exaltata Mahana Preserved_Specimen Munro, G.C. 178 Lanai BISH 1913-10-01
62 Nephrolepis exaltata Punaluu-Kaipapau Preserved_Specimen Forbes, C.N. Oahu BISH 1909-05-08
63 Nephrolepis exaltata near Puu Kana, Waianae Mts Preserved_Specimen Gillespie, J.W. 1062 Oahu BISH 1927-07-06
64 Nephrolepis exaltata Koolau Mts., Puu Peahinaia, Paalaa-Kawailoa divide Preserved_Specimen St.John, H. 11121 Oahu BISH 1931-10-04
65 Nephrolepis exaltata Mahana Preserved_Specimen Munro, G.C. 182 Lanai BISH 1913-10-01
66 Nephrolepis exaltata Waianae Mts., Kunia trail just E of Puu Kanehoa Preserved_Specimen Stone, B.C. 3312 Oahu BISH 1960-03-26
67 Nephrolepis exaltata Laie, behind CCH Preserved_Specimen Leong, S. 12 Oahu BISH 1960-10-18
68 Nephrolepis exaltata Molokai Preserved_Specimen Stokes, J.F.G. 10 Molokai BISH 1909-01-01
69 Nephrolepis exaltata Kilauea, Ka Loko Reservoir Preserved_Specimen Forbes, C.N. 589 Kauai BISH 1916-10-13
70 Nephrolepis exaltata Kaawaloa, Hawaii Preserved_Specimen Souza, W. Hawaii BISH 1984-10-04
71 Nephrolepis exaltata Waiamao Hill near Waiohinu, Kau Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-07-28
72 Nephrolepis exaltata peak of Mt. Olympus Preserved_Specimen Garber, D.W. 293 Oahu BISH 1920-03-07
73 Nephrolepis exaltata East Maui, Koolau Gap, E side Preserved_Specimen Hobdy, R.W. 3383 Maui BISH 1991-07-18
74 Nephrolepis exaltata Kalaupapa Trail Preserved_Specimen St.John, H. 25255 Molokai BISH 1953-12-25
75 Nephrolepis exaltata Koolau Mts., Puu Uau, Keaiwa Heiau Preserved_Specimen Krajina, V.J. 611104001 Oahu BISH 1961-11-04
76 Nephrolepis exaltata East Maui, Koolau Gap, E side Preserved_Specimen Hobdy, R.W. 3383 Maui BISH 1991-07-18
77 Nephrolepis exaltata East Maui, Hana District, SW side of Haleakala, Lualailua Preserved_Specimen Forbes, C.N. 2108 Maui BISH 1920-03-25
78 Nephrolepis exaltata Pauoa Valley Preserved_Specimen Mann, H. 142393 Oahu BISH 1864-01-01
79 Nephrolepis exaltata Hulopoe Gulch, head of Preserved_Specimen Degener, O. 28525 Lanai BISH 1963-08-09
80 Nephrolepis exaltata Oahu Preserved_Specimen Rémy, J. 43 Oahu BISH 1851-01-01
81 Nephrolepis exaltata Haena, Waikapalae Cave Preserved_Specimen Chock, A.K. 408 Kauai BISH 1952-02-07
82 Nephrolepis exaltata Waianae Mts., near Kaala Preserved_Specimen Selling, O. 3553 Oahu BISH 1938-09-25
83 Nephrolepis exaltata Mts. behind Honolulu Preserved_Specimen U.S. Exploring Expedition 5 Oahu BISH 1838-01-01
84 Nephrolepis exaltata Palolo. Preserved_Specimen Lyon, H.L. 1015 Oahu BISH 1907-11-17
85 Nephrolepis exaltata N. Kona, Holualoa FR, S slope of Hualalai Preserved_Specimen Kondo, Y. Hawaii BISH 1946-07-13
86 Nephrolepis exaltata near Hilo Preserved_Specimen U.S. Exploring Expedition Hawaii BISH 1838-01-01
87 Nephrolepis exaltata Kilauea, floor of Kilaueaiki Preserved_Specimen Neal, M.C. Hawaii BISH 1929-08-03
88 Nephrolepis exaltata Waiamao Hill, Waiohinu, Kau Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-07-28
89 Nephrolepis exaltata Kaimuki Preserved_Specimen Fullaway, D. Oahu BISH 1920-01-01
90 Nephrolepis exaltata S ridge of Kipapa Gulch, Waipio, Koolau Mts Preserved_Specimen Bryan Jr., E.H. Oahu BISH 1932-05-15
91 Nephrolepis exaltata East Maui, upper ditch trail. Preserved_Specimen Lyon, H.L. Maui BISH 1908-11-23
92 Nephrolepis exaltata Honolulu Preserved_Specimen Pukui, M.K. Oahu BISH 1931-01-01
93 Nephrolepis exaltata Kaala, Waianae Mts Preserved_Specimen Swezey, O.H. Oahu BISH 1932-10-23
94 Nephrolepis exaltata Molokai Preserved_Specimen Stokes, J.F.G. Molokai BISH 1909-01-01
95 Nephrolepis exaltata Palolo Valley Preserved_Specimen Forbes, C.N. Oahu BISH 1908-11-07
96 Nephrolepis exaltata Pacific Heights Preserved_Specimen Bryan, W.A. Oahu BISH 1903-12-20
97 Nephrolepis exaltata Kaali Preserved_Specimen Stokes, J.F.G. Niihau BISH 1912-01-01
98 Nephrolepis exaltata Punaluu-Kaipapau, Koolau Mts Preserved_Specimen Forbes, C.N. Oahu BISH 1907-05-03
99 Nephrolepis exaltata Waikane-Schofield Trail. Preserved_Specimen Lohman, M.L. Oahu BISH 1947-09-20
100 Nephrolepis exaltata Kau Rd. below 1/2 way house Preserved_Specimen Giffard, W.M. Hawaii BISH 1927-07-11
101 Nephrolepis exaltata Molokai Preserved_Specimen Stokes, J.F.G. Molokai BISH 1909-01-01
102 Nephrolepis exaltata Lihue, N side Preserved_Specimen Forbes, C.N. 38 Kauai BISH 1909-07-10
103 Nephrolepis exaltata Pupukea Summit, Waialua, Kahuku Divide Preserved_Specimen Barrere, D. Oahu BISH 1959-12-13
104 Nephrolepis exaltata Pukoo Ridge Preserved_Specimen Forbes, C.N. 421 Molokai BISH 1912-08-01
105 Nephrolepis exaltata Bryan's Mt. House Trail at Grant Boundary Preserved_Specimen Krajina, V.J. 620520031 Oahu BISH 1962-05-20
106 Nephrolepis exaltata spur ridge off rd. from Lihue to Lawai near Koloa Jct Preserved_Specimen Stone, B.C. 737 Kauai BISH 1955-06-15
107 Nephrolepis exaltata Preserved_Specimen BISH
108 Nephrolepis exaltata main ridge SW from Puu Lanihuli, Kalihi-Nuuanu Preserved_Specimen St.John, H. 11187 Oahu BISH 1931-11-29
109 Nephrolepis exaltata No locality information available HUMAN_OBSERVATION 1441951053 Oahu HAPI 12/22/1997
110 Nephrolepis exaltata Hanalei, Limahuli Valley forest along stream; with Psidium, Aleurites, Schefflera, Metrosideros, Pisonia, Hedyotis. On guava Preserved_Specimen Flynn, T.W. 2310 Kauai BISH 1987-08-20
111 Nephrolepis exaltata Koolau Mountain Range: Kalauao ridge, Ewa Forest Reserve thicket below forest Preserved_Specimen St.John, H. 12989 Oahu BISH 1933-03-29
112 Nephrolepis exaltata Waipio Trail, Hamakua on mossy rock ledge Preserved_Specimen St.John, H. 24001 Hawaii BISH 1950-01-01
113 Nephrolepis exaltata Waianae Mts., Halona, Lualualei Forest Reserve; 2nd gulch S of Pohakea Pass, steep grassy slope Preserved_Specimen St.John, H. 13146 Oahu BISH 1933-05-12
114 Nephrolepis exaltata National Park, Kipuka Puaulu, ~1/2 way around nature trail Acacia, Sapindus, Microlepia Preserved_Specimen Newell, C.L. 1039 Hawaii BISH 1967-06-28
115 Nephrolepis exaltata Koolau Mts., Maakua-Papali ridge rain forest, on ground Preserved_Specimen Wilson, K.A. 152 Oahu BISH 1952-10-26
116 Nephrolepis exaltata W Maui, Lahaina District, Lihau Lowland Koa Mesic Forest on a ridge at 3300 ft beneath a canopy of Acacia koa, Dodonaea, Coprosma and Myrsine with a understory of Diplazium sandwichianum, Doodia kunthiana, Microlepia, and Carex meyenii Preserved_Specimen Welton, P. 1005 Maui BISH 1991-07-09
117 Nephrolepis exaltata Oahu Forest National Wildlife Refuge, Kipapa Trail, ~1.75 miles from refuge boundary, north side of trail. Metrosideros-Acacia-Dicranopteris wet lowland forest invaded by Psidium cattleianum at colleciton site. Preserved_Specimen Reynolds, L.S. 195 Oahu BISH 2006-07-05
118 Nephrolepis exaltata Nuuanu, Southern Koolau Mountains. On switchback trail to Konahuanui. Growing on side of trail in wet forest dominated by Metrosideros tremuloides, M. polymorpha, Kadua affinis, Machaerina angustifolia, Wikstroemia oahuensis, Dicranopteris linearis, and others. Preserved_Specimen Thomas, M.K. 352 Oahu BISH 2022-04-03
119 Nephrolepis exaltata NW side Ulumawai, Kailua dry, rocky slide Preserved_Specimen Fosberg, F.R. 13551 Oahu BISH 1937-01-24
120 Nephrolepis exaltata Poamoho near cabin Growing along trail in Metrosideros-Dicranopteris-Cibotium dominated wet forest. Growing with N. cordifolia. Preserved_Specimen Thomas, M.K. 103 Oahu BISH 2021-08-18
121 Nephrolepis exaltata N slope Puu Olomana, Kailua foot of precipice in brush, moist, shady Preserved_Specimen Fosberg, F.R. 9024 Oahu BISH 1932-11-13
122 Nephrolepis exaltata Chain of Craters Rd. ~10.4 mi from intersection with Crater Rim Rd.; NW of hwy. ~35 m Metrosideros, Andropogon Preserved_Specimen Newell, C.L. 815 Hawaii BISH 1967-01-23
123 Nephrolepis exaltata Kulani Prison Rd. ~5.4 mi from intersection with Kona-Hilo Hwy.; ~20 m N of road Cibotium forest with scattered Metrosideros Preserved_Specimen Newell, C.L. 904 Hawaii BISH 1967-01-25
124 Nephrolepis exaltata W side of Kahana Valley, 6 km upstream moist, open, fern covered ridge Preserved_Specimen Fosberg, F.R. 13729 Oahu BISH 1937-04-18
125 Nephrolepis exaltata below Puhima Crater edges of hot spot; barren fumarole Preserved_Specimen Fosberg, F.R. 60567 Hawaii BISH 1980-09-07
126 Nephrolepis exaltata W slope of Wailupe Gulch base of rock outcrop Preserved_Specimen Egler, F.E. 37289 Oahu BISH 1937-03-02
127 Nephrolepis exaltata Mokuleia trail. Pahole NAR. Northern Waianae mountains. Growing along trail in mesic dry forest dominated by Psidium cattleianum, Schinus terebinthifolius, Leucobryum gracile, Pyrrhobryum spiniforme, and Christella parasitica. Also growing with Microlepia setosa, Alyxia stellata, and Metrosideros polymorpha. Preserved_Specimen Thomas, M.K. 632 Oahu BISH 2024-02-29
128 Nephrolepis exaltata Waianae Mts., Waianae Kai Trail, left side of trail on way up , above junction Alien mixed mesic forest with Kukui (Aleurites moluccana), Guava (Psidium cattleianum), and Silk oak (Grevillea robusta). Preserved_Specimen Crago, L.M. 2004001 Oahu BISH 2004-11-09
129 Nephrolepis exaltata Hawaii, Volcano Agricultural Experiment Station Farm, N. of Kilauea. In dense Cibotium glaucum forest with emergent Metrosideros. On branch of mossy tree. Preserved_Specimen Fosberg, F.R. 65387 Hawaii BISH 1988-01-30
130 Nephrolepis exaltata upper Lihue ditch rain forest Preserved_Specimen MacDaniels, L.H. 822 Kauai BISH 1927-02-21
131 Nephrolepis exaltata end of Hilina Pali Rd.; down trail ~1.5 mi open Metrosideros, Diospyros forest on aa lava Preserved_Specimen Newell, C.L. 267 Hawaii BISH 1966-08-02
132 Nephrolepis exaltata Hilo forest Preserved_Specimen Newell, Brother M. Hawaii BISH 1910-01-01
133 Nephrolepis exaltata Waimano Trail moist back Preserved_Specimen Bryan Jr., E.H. 1550 Oahu BISH 1941-03-23
134 Nephrolepis exaltata Pupukea Ridge sandy loam Preserved_Specimen Sanchez, S. 26 Oahu BISH 1986-01-21
135 Nephrolepis exaltata Aiea Loop Trail, Koolau Mts on bank in shade among Nephrolepis hirsutula Preserved_Specimen Yee, R.S.N. 9 Oahu BISH 1971-08-12
136 Nephrolepis exaltata near Hilo terminal portion of 1880-1 lava flow Preserved_Specimen Forbes, C.N. 553 Hawaii BISH 1915-05-17
137 Nephrolepis exaltata Koaie Canyon diverse mesic forest; small stream drainage near base of N facing canyon wall; Dodonaea, Metrosideros, Pleomele, Perrottetia, Diospyros sandwicensis, Psidium cattleianum, Boehmeria, Lantana, Kalanchoe pinnata, Rubus rosifolius, Christella dentata; rocks along stream Preserved_Specimen Trauernicht, C. 714 Kauai BISH 2009-02-02
138 Nephrolepis exaltata gully near Haleakala choking dry stream bed; with Sapindus oahuensis, Rauvolfia sandwicensis, Antidesma pulvinatum Preserved_Specimen Takeuchi, W.N. 3310 Maui BISH 1987-01-01
139 Nephrolepis exaltata Kapano Gulch, Kalulu woods Preserved_Specimen St.John, H. 18769 Lanai BISH 1938-04-10
140 Nephrolepis exaltata N-S Halawa ridge, Koolau Mts lower forest, koa zone Preserved_Specimen St.John, H. 20383 Oahu BISH 1943-12-12
141 Nephrolepis exaltata below Ewa and Sisal coral plains, in caves Preserved_Specimen Forbes, C.N. 2324 Oahu BISH 1916-11-14
142 Nephrolepis exaltata Kipapa Gulch, Waipio Metrosideros-Antidesma forest Preserved_Specimen Grant, M.L. 7168 Oahu BISH 1934-08-06
143 Nephrolepis exaltata Kulani Prison Rd. ~7.4 mi from intersection with Kona-Hilo Hwy.; ~50 m L of road Metrosideros, Cibotium, Pritchardia Preserved_Specimen Newell, C.L. 310 Hawaii BISH 1966-08-09
144 Nephrolepis exaltata Kulanapahu Cone, Ulapalakua rock crevices on broken lava ridges Preserved_Specimen Fosberg, F.R. 48363 Maui BISH 1967-03-25
145 Nephrolepis exaltata Kau, Moaula open, dry Preserved_Specimen Russ, G.W. 35 Hawaii BISH 1927-06-01
146 Nephrolepis exaltata Leaukahi, Koloa open woods Preserved_Specimen St.John, H. 13449 Kauai BISH 1933-12-22
147 Nephrolepis exaltata Haena, Waikapalae Cave on rock in cave above water Preserved_Specimen Chock, A.K. 411 Kauai BISH 1952-02-07
148 Nephrolepis exaltata 16.2 mi W of Hilo on Saddle Rd Metrosideros and Cibotium forest on lava bed; dark Preserved_Specimen Stuessy, T.F. 6124 Hawaii BISH 1982-05-30
149 Nephrolepis exaltata W ridge of Honomuni moist Metrosideros forest Preserved_Specimen St.John, H. 25199 Molokai BISH 1953-12-23
150 Nephrolepis exaltata Kaali Cliff, first valley W On face of overhanging cliff Preserved_Specimen St.John, H. 22844 Niihau BISH 1947-08-16
151 Nephrolepis exaltata W side of Kahana Valley, 6 km upstream moist, open, fern covered ridge Preserved_Specimen Fosberg, F.R. 13729 Oahu BISH 1937-04-18
152 Nephrolepis exaltata Makaleha Valley woods Preserved_Specimen Smith, F.G. 3 Oahu BISH 1929-01-14
153 Nephrolepis exaltata SW flank of Mauna Loa, Manuka in kipuka; closed Metrosideros forest with Acacia, Cibotium, Coprosma, Myrsine; on soil and aa Preserved_Specimen Higashino, P.K. 6229 Hawaii BISH 1977-07-21
154 Nephrolepis exaltata Kapua with Lantana, Dracaena; gulch Preserved_Specimen Bryan Jr., E.H. Hawaii BISH 1932-11-24
155 Nephrolepis exaltata Kaala, Waianae Mts valley bottom Preserved_Specimen Bryan Jr., E.H. Oahu BISH 1932-10-23
156 Nephrolepis exaltata forest reserve moist shaded forest floor along jeep road Preserved_Specimen Herbst, D.R. 5107 Molokai BISH 1974-11-13
157 Nephrolepis exaltata Lyon Arboretum, nose of ridge above section 10 on koa Preserved_Specimen Baker, R.F. 128 Oahu BISH 1975-06-13
158 Nephrolepis exaltata S. Opaeula Gulch, Paalaa wooded ridge Preserved_Specimen St.John, H. 10624 Oahu BISH 1930-11-09
159 Nephrolepis exaltata Waianae Mts., Makaha Valley rain forest, on ground Preserved_Specimen Wilson, K.A. 192 Oahu BISH 1952-11-30
160 Nephrolepis exaltata Haena, Waikapalae Cave on rock in cave above water Preserved_Specimen Chock, A.K. 411 Kauai BISH 1952-02-07
161 Nephrolepis exaltata 2nd gulch E of Puu Kaupakuhale, Mokuleia, Kaala, Waianae Mts wooded ridge Preserved_Specimen Fosberg, F.R. 8916 Oahu BISH 1932-10-23
162 Nephrolepis exaltata Puu Kalena, Waianae Mts., Waianae Kai woods Preserved_Specimen St.John, H. 12948 Oahu BISH 1933-03-12
163 Nephrolepis exaltata Laie-Malaekahana ridge, Koolau Mts woods Preserved_Specimen St.John, H. 13056 Oahu BISH 1933-04-15
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