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Centella asiatica

(L.) Urb. (1879)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Apiales Family: Apiaceae Genus: Centella

asiatic pennywort, gotu kola, gotukola, pohe kula

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Description

Key Characters:

Growth Form: Caulescent perennial herbs.

Stems: Caulescent, stems 1 to several, creeping, rooting at the nodes, glabrous to flocose and glabrate.

Roots:

Leaves: Leaves simple. Alternate. Blades orbicular–reniform to ovate–cordate, 2–6 cm long, 1.5–5 cm wide. Surfaces glabrous. Margins crenate to entire. Palmately-veined; 5–9 veined. Sheathing petioles; 3–30 cm long. Stipules absent.

Flowers: Flowers 2–3 in umbels. Peduncles 1.2–4 cm long, Involucral bracts usually 2, ovate to suborbicular, 2.5–3 mm long, 1–5–2.5 mm wide, scariously margined, pedicels short, 0–4 mm long. Flowers bisexual (perfect). Calyx teeth obscure or absent. Corolla of 5 petals; petals white to purplish or rose. Stamens 5, inserted on an epigynous disk. Ovary inferior, 2-celled; ovules 1 per cells, anatropous; styles 2, usually swollen at base into a stylopodium. Styles short, the stylopodium absent, carpophore entire.

Fruit: Fruit ovoid to orbicular; 2–5 mm in diameter; mericarps strongly flattened laterally; ribs usually 7–9; connected by conspicuous reticulations; an oil–bearing layer beneath the outer pericarp; but companion canals and vittae are usually absent. Seeds 1 per mericarp; embryo small; endosperm cartilaginous. seed face plane.

Ploidy: 2n = 17; 18*; 19; 20; 22; 33

Habitat: Naturalized in a wide range of habitats including meadows; ditches; bogs; and wet forest.

Elevation Range: 0–1,000 m.

Historical Distribution

Images

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Synonyms (39)

Centella asiatica f. crispata (Maxim.) H.Hara (1954)
Centella asiatica var. boninensis (Nakai ex Tuyama) H.Hara (1954)
Centella asiatica var. crispata (Maxim.) Y.Yabe (1902)
Centella asiatica var. cristata Makino (1904)
Centella boninensis Nakai ex Tuyama (1936)
Centella hirtella Nannf. (1924)
Centella ulugurensis (Engl.) Domin (1908)
Chondrocarpus asiaticus Nutt. (1818)
Chondrocarpus triflorus Nutt. (1818)
Glyceria asiatica Nutt. (1818)
Glyceria triflora Nutt. (1818)
Hydrocotyle abbreviata A.Rich. (1820)
Hydrocotyle abyssinica Gand. (1918)
Hydrocotyle artensis Montrouz. (1860)
Hydrocotyle asiatica L. (1753)
Hydrocotyle asiatica var. crispata Maxim. (1886)
Hydrocotyle asiatica var. monantha F.Muell. (1874)
Hydrocotyle brasiliensis Scheidw. (1842)
Hydrocotyle brevipedata St.-Lag. (1880)
Hydrocotyle brevipes DC. (1830)
Hydrocotyle dentata A.Rich. (1820)
Hydrocotyle ficarifolia Stokes (1812)
Hydrocotyle ficarioides Lam. (1789)
Hydrocotyle hebecarpa DC. (1830)
Hydrocotyle inaequipes DC. (1830)
Hydrocotyle indivisa Banks & Sol. ex Hook.f. (1852)
Hydrocotyle leptostachys Spreng. (1824)
Hydrocotyle lunata Lam. (1789)
Hydrocotyle lurida Hance (1852)
Hydrocotyle natans Thunb. (1798)
Hydrocotyle nummularioides A.Rich. (1820)
Hydrocotyle pallida DC. (1830)
Hydrocotyle reniformis Walter (1788)
Hydrocotyle sarmentosa Salisb. (1796)
Hydrocotyle sylvicola Cordem. (1895)
Hydrocotyle thunbergiana Spreng. (1820)
Hydrocotyle ulugurensis Engl. (1900)
Hydrocotyle wightiana Wall. (1832)
Trisanthus cochinchinensis Lour. (1790)

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Lana'i Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents

Water/Ocean

Vegetative Reproduction

Pollinators

Notes

  • Description digitized by Pumehana Imada
  • Native to Asia. Naturalized prior to 1871 (Hillebrand, 1888). The plant widespread in the Hawaiian Islands is apparently the ubiquitous Asian diploid of this polyploid complex.

Bibliography

Name Published In: C.F.P.von Martius & auct. suc. (eds.), Fl. Bras. 11(1): 287 (1879)

Other References

Wagner et al. 1990:201 (K, O, Mo, L, M, H); Staples & Herbst 2005:112 (KEY), 115 (DESCR)

Wagner, W. L., Herbst, D. R., & Sohmer, S. H. (1999). Manual of the Flowering Plants of Hawai'i, Vols. 1 and 2 (No. Edn 2). University of Hawai'i and Bishop Museum Press.

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Centella asiatica near the U.S. Forest Service Experimental Tree Planting Site within Halepuaa Forest Reserve in the Puna District Preserved_Specimen Clarke, G. 79.217 Hawaii BISH 1979-03-08
2 Centella asiatica Kaneohe 47-775 Ahilama Rd. (private land holding) Preserved_Specimen Annable, C.R. 3361 Oahu BISH 1997-03-15
3 Centella asiatica ahupuaa of Halepuaa in the Puna District, in the area of teh experimental tree planting site on the Puna Trail OBSERVATION Curtis Daehler Hawaii HAPI 7/19/2007
4 Centella asiatica Oahu Is., koolau range, poamoho trail OBSERVATION Curtis Daehler Oahu HAPI 3/18/2007
5 Centella asiatica Makawaon suojelualue, Kula, HI, US OBSERVATION Curtis Daehler Hawaii HAPI 3/18/2007
6 Centella asiatica Kailua, Ulumawao Ridge Somewhat common on ridges, beneath Osteomeles. Preserved_Specimen Davis, J. 90 Oahu BISH 1979-03-07
7 Centella asiatica Windward side of island, Anapuhi to Wainene, long coastal boulder-strewn beach with vertical cliff at back 10--30 m from water Cliff with Artemisia australis, Chenopodium oahuense, Schiedea globosa, Kalanchoe pinnata, Tetramolopium sylvae, Lythrum maritimum, and Hedyotis littoralis, among others. Preserved_Specimen Wagner, W.L. 5746 Molokai BISH 1987-08-01
8 Centella asiatica Waianae Mts., Puu Manawahua Common ground cover in woods. Preserved_Specimen St.John, H. 9896 Oahu BISH 1929-09-29
9 Centella asiatica Kalepa Ridge Windward side of northern part of ridge. Preserved_Specimen Corn, C. Kauai BISH 1977-07-20
10 Centella asiatica Lanai City, S section Weed in waste area. Preserved_Specimen Nagata, K.M. 3493 Lanai BISH 1986-06-02
11 Centella asiatica East Maui, Hana District, Waihoi Valley Found in marshy depression; seemed to occur in wet disturbed areas with several introduced grasses and sedges. Preserved_Specimen Harrison, B. 25 Maui BISH 1972-09-21
12 Centella asiatica Hilo District, Onomea Valley, Hawaii Tropical Botanical Garden grounds Creeping on ground in deep shade beneath coconut palms, Pandanus trees, and Alexander palms; exposed to salt spray. Preserved_Specimen Staples, G.W. 529 Hawaii BISH 1990-01-08
13 Centella asiatica [Hawaiian Islands; U.S.A.; Molokai; Hawaii; Maui: Molokai: Lanikula Anele was collected at Lanikula under the kukui trees. There were many kukui seedlings, noni seedlings, wiki, and kukae-pua`a among other plants in the beautiful kukui grove. Preserved_Specimen Funk, E.J. 920052 Molokai BISH 1992-05-03
14 Centella asiatica Koolau Mts., Pupukea - along summit trail On grassy summit. Observed several Pritchardia stands. Preserved_Specimen Stone, B.C. 2815 Oahu BISH 1959-05-04
15 Centella asiatica Between Waiahole and Waikane valleys, windward side of Koolau Range On wind-swept ridge. Preserved_Specimen Harris, J.A. 242047 Oahu BISH 1924-08-16
16 Centella asiatica ahupuaa of Halepuaa in the Puna District, in the area of teh experimental tree planting site on the Puna Trail The soil in the area was crushed lava or cinder. The plant was under tall grasses that shielded it from the direct sunlight. Preserved_Specimen Yoshida, L. 79.066 Hawaii BISH 1979-03-08
17 Centella asiatica Waikane-Schofield Trail, Koolau Mts., Waikane Wet trail-side bank. Preserved_Specimen Fosberg, F.R. 8783 Oahu BISH 1932-10-16
18 Centella asiatica Wailua Game Reserve, above Wailua Vegetation on plots stripmined for bauxite ore in 1958-60. Tree plot area. Preserved_Specimen Howard, R.A. 20251 Kauai BISH 1985-10-20
19 Centella asiatica Koloa District. Lower slope of Mt. Kahili off the TV Relay-Alexander Dam Rd., ca. 1.5 mi above the Wahiawa Bog Associated with Hibiscus kahilii, Clermontia clermontioides, Psychotria, Dubautia, Psidium, Metrosideros polymorpha, etc. Preserved_Specimen Gustafson, R. 1056 Kauai BISH 1978-11-25
20 Centella asiatica Waikiki District, Honolulu Vacant lot. Preserved_Specimen Baker, R.J. 228 Oahu BISH 1932-07-20
21 Centella asiatica East Maui, Waihoi Valley, Transect 1 In wet open disturbed areas with Paspalum conjugatum. Preserved_Specimen Harrison, B. 20 Maui BISH 1972-06-15
22 Centella asiatica Kipapa Trail, ~100 m from refuge boundary, along edges of trail wet lowland forest; Metrosideros, Acacia, Dicranopteris Preserved_Specimen Reynolds, L.S. 72 Oahu BISH 2006-05-31
23 Centella asiatica Lyon Arboretum, Honolulu; Upper Lawn Makai Around base of palms near gate. Very common weed. Wide range of habitats. Preserved_Specimen Baker, R.F. 81 Oahu BISH 1975-04-25
24 Centella asiatica Near mouth of Waipio Stream, Waipio Valley, Big Island, Hawaii Growing on edge of outflow to the stream, from Cyperus papyrus pond area, in sandy wet soil. Preserved_Specimen Crago, L.M. 2005115 Hawaii BISH 2005-03-19
25 Centella asiatica Wailau Valley, n. coast East Molokai Rocky flat above tide level. Preserved_Specimen Fosberg, F.R. 9649 Molokai BISH 1933-07-04
26 Centella asiatica Mokuauia Island, Laie In crevices in coral rock. Preserved_Specimen Fosberg, F.R. 14169 Oahu BISH 1937-07-04
27 Centella asiatica Tantalus, Koolau Mts Wet forest, along trail - disturbed ground. Preserved_Specimen Stemmermann, R.L. 426 Oahu BISH 1973-08-09
28 Centella asiatica Located down Kona-Hilo Hwy 4 mi past Hongo Store towards Hilo; turn right onto Fern Estates Road. Plant was about 200 m down Fern Estates Road, on right side. (plot 55) Scrub forest community. Preserved_Specimen Newell, C.L. 671 Hawaii BISH 1966-08-31
29 Centella asiatica Honolulu. Crest of Koolau Range, near Puu Konahuanui, ca. 3.5 air mi NNE of Univ. of Hawaii campus Native forest; along trail. Preserved_Specimen Young, D.A. 1639 Oahu BISH 1979-01-18
30 Centella asiatica Honolulu: Manoa Valley; Lyon Arboretum-Middle Lawn Makai Collected under eaves of new Greenhouse. Preserved_Specimen Nagata, K.M. 439 Oahu BISH 1968-06-28
31 Centella asiatica Hawaii National Park, Kilauea, Crater Rim Road, near Fern Jungle Trail Locally abundant. Preserved_Specimen Fagerlund, G.O. 1080 Hawaii BISH 1945-06-13
32 Centella asiatica Ditch Trail Kahana Valley, Koolau Mts. Kahana Wet forest, hanging over bank along tract. Preserved_Specimen Fosberg, F.R. 10410 Oahu BISH 1933-11-26
33 Centella asiatica Along Schofield-Waikane Trail, leeward of Koolau Ridge Abundant on moist roadside bank and along trail. Preserved_Specimen Crosby, M.R. 1594 Oahu BISH 1964-06-30
34 Centella asiatica Koolau Range, Manoa; Ridge trail back of Woodlawn Growing on side of tree. Preserved_Specimen Kuykendall, J.R. 66 Oahu BISH 1942-12-31
35 Centella asiatica Lyon Arboretum, Honolulu; Upper Lawn Makai Around base of palms near gate. Very common weed. Preserved_Specimen Baker, R.F. 81 Oahu BISH 1975-04-25
36 Centella asiatica Makawao District, Koolau Forest Reserve, N slope of Haleakala. Small gulch below dilapidated flume, 0.2 mi E of Waikamoi Stream Muddy streambed in wet forest. Preserved_Specimen Lammers, T.G. 5655 Maui BISH 1984-07-13
37 Centella asiatica Lanai City Weed in a lawn. Preserved_Specimen Hobdy, R.W. 2547 Lanai BISH 1986-03-09
38 Centella asiatica So. ridge So. Opaeula Gulch, Koolau Mts. Paalaa Roadside mud. Preserved_Specimen Fosberg, F.R. 8738 Oahu BISH 1932-09-25
39 Centella asiatica Pupukea Forest, Koolauloa Creek bed Aleurites Forest. Preserved_Specimen Grant, M.L. 7154 Oahu BISH 1934-08-01
40 Centella asiatica Koloa District, Lawai Valley, National Tropical Botanical Garden Common in moist, shady areas of valley. Preserved_Specimen Herbst, D.R. 2286 Kauai BISH 1972-01-03
41 Centella asiatica Nuuanu Water Shed, site of ruins of King Kamehameha III's summer home Preserved_Specimen Mill, S.W. 13 Oahu BISH 1983-08-05
42 Centella asiatica Aiea In pineapple field. Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
43 Centella asiatica Waianae Ridge, Maunauna, SE slope In Acacia koa forest. Preserved_Specimen van Royen, P. 10126 Oahu BISH 1967-07-04
44 Centella asiatica Lahaina District, Road to Hailau [probably] on Kaulalewelewe between Honokowai and Mahinahina Valleys Disturbed mesic forest Preserved_Specimen Ray, G. 1062 Maui BISH 1999-02-18
45 Centella asiatica open wet forest Preserved_Specimen Allison, A. Oahu BISH 2021-01-17
46 Centella asiatica Top of pali overlooking leper settlement, Kalaupapa Shade of bush. Preserved_Specimen St.John, H. 12540 Molokai BISH 1932-12-24
47 Centella asiatica East Maui, Hana Rain Forest Project In open bog with Carex svenonis and Erechtites valerianifolia. Preserved_Specimen Harrison, B. 316 Maui BISH 1973-07-09
48 Centella asiatica Kalaupapa; Papapaiki, coast Preserved_Specimen Wysong, M.L. 581 Molokai BISH 2005-01-12
49 Centella asiatica Honolulu, Manoa Valley, Manoa Arboretum Preserved_Specimen Russell, P. 23 Oahu BISH 1932-07-05
50 Centella asiatica Honolulu, Bishop Museum campus Preserved_Specimen Brown, F.B.H. 1323 Oahu BISH 1925-01-03
51 Centella asiatica Nuuanu Valley Preserved_Specimen Forbes, C.N. 1421 Oahu BISH 1909-12-12
52 Centella asiatica West of Waialua, near irrigation ditch, edge of cane field Preserved_Specimen Kerr, M. Oahu BISH 1946-06-30
53 Centella asiatica Manoa Arboretum Preserved_Specimen Caum, E.L. Oahu BISH 1938-07-07
54 Centella asiatica Mokolii Preserved_Specimen Neal, M.C. Oahu BISH 1938-09-24
55 Centella asiatica near the U.S. Forest Service Experimental Tree Planting Site within Halepuaa Forest Reserve in the Puna District Preserved_Specimen Clarke, G. 79.217 Hawaii BISH 1979-03-08
56 Centella asiatica Kaneohe 47-775 Ahilama Rd. (private land holding) Preserved_Specimen Annable, C.R. 3361 Oahu BISH 1997-03-15
57 Centella asiatica ahupuaa of Halepuaa in the Puna District, in the area of teh experimental tree planting site on the Puna Trail OBSERVATION Curtis Daehler Hawaii HAPI 7/19/2007
58 Centella asiatica Oahu Is., koolau range, poamoho trail OBSERVATION Curtis Daehler Oahu HAPI 3/18/2007
59 Centella asiatica Makawaon suojelualue, Kula, HI, US OBSERVATION Curtis Daehler Hawaii HAPI 3/18/2007
60 Centella asiatica Kailua, Ulumawao Ridge Somewhat common on ridges, beneath Osteomeles. Preserved_Specimen Davis, J. 90 Oahu BISH 1979-03-07
61 Centella asiatica Windward side of island, Anapuhi to Wainene, long coastal boulder-strewn beach with vertical cliff at back 10--30 m from water Cliff with Artemisia australis, Chenopodium oahuense, Schiedea globosa, Kalanchoe pinnata, Tetramolopium sylvae, Lythrum maritimum, and Hedyotis littoralis, among others. Preserved_Specimen Wagner, W.L. 5746 Molokai BISH 1987-08-01
62 Centella asiatica Waianae Mts., Puu Manawahua Common ground cover in woods. Preserved_Specimen St.John, H. 9896 Oahu BISH 1929-09-29
63 Centella asiatica Kalepa Ridge Windward side of northern part of ridge. Preserved_Specimen Corn, C. Kauai BISH 1977-07-20
64 Centella asiatica Lanai City, S section Weed in waste area. Preserved_Specimen Nagata, K.M. 3493 Lanai BISH 1986-06-02
65 Centella asiatica East Maui, Hana District, Waihoi Valley Found in marshy depression; seemed to occur in wet disturbed areas with several introduced grasses and sedges. Preserved_Specimen Harrison, B. 25 Maui BISH 1972-09-21
66 Centella asiatica Hilo District, Onomea Valley, Hawaii Tropical Botanical Garden grounds Creeping on ground in deep shade beneath coconut palms, Pandanus trees, and Alexander palms; exposed to salt spray. Preserved_Specimen Staples, G.W. 529 Hawaii BISH 1990-01-08
67 Centella asiatica [Hawaiian Islands; U.S.A.; Molokai; Hawaii; Maui: Molokai: Lanikula Anele was collected at Lanikula under the kukui trees. There were many kukui seedlings, noni seedlings, wiki, and kukae-pua`a among other plants in the beautiful kukui grove. Preserved_Specimen Funk, E.J. 920052 Molokai BISH 1992-05-03
68 Centella asiatica Koolau Mts., Pupukea - along summit trail On grassy summit. Observed several Pritchardia stands. Preserved_Specimen Stone, B.C. 2815 Oahu BISH 1959-05-04
69 Centella asiatica Between Waiahole and Waikane valleys, windward side of Koolau Range On wind-swept ridge. Preserved_Specimen Harris, J.A. 242047 Oahu BISH 1924-08-16
70 Centella asiatica ahupuaa of Halepuaa in the Puna District, in the area of teh experimental tree planting site on the Puna Trail The soil in the area was crushed lava or cinder. The plant was under tall grasses that shielded it from the direct sunlight. Preserved_Specimen Yoshida, L. 79.066 Hawaii BISH 1979-03-08
71 Centella asiatica Waikane-Schofield Trail, Koolau Mts., Waikane Wet trail-side bank. Preserved_Specimen Fosberg, F.R. 8783 Oahu BISH 1932-10-16
72 Centella asiatica Wailua Game Reserve, above Wailua Vegetation on plots stripmined for bauxite ore in 1958-60. Tree plot area. Preserved_Specimen Howard, R.A. 20251 Kauai BISH 1985-10-20
73 Centella asiatica Koloa District. Lower slope of Mt. Kahili off the TV Relay-Alexander Dam Rd., ca. 1.5 mi above the Wahiawa Bog Associated with Hibiscus kahilii, Clermontia clermontioides, Psychotria, Dubautia, Psidium, Metrosideros polymorpha, etc. Preserved_Specimen Gustafson, R. 1056 Kauai BISH 1978-11-25
74 Centella asiatica Waikiki District, Honolulu Vacant lot. Preserved_Specimen Baker, R.J. 228 Oahu BISH 1932-07-20
75 Centella asiatica East Maui, Waihoi Valley, Transect 1 In wet open disturbed areas with Paspalum conjugatum. Preserved_Specimen Harrison, B. 20 Maui BISH 1972-06-15
76 Centella asiatica Kipapa Trail, ~100 m from refuge boundary, along edges of trail wet lowland forest; Metrosideros, Acacia, Dicranopteris Preserved_Specimen Reynolds, L.S. 72 Oahu BISH 2006-05-31
77 Centella asiatica Lyon Arboretum, Honolulu; Upper Lawn Makai Around base of palms near gate. Very common weed. Wide range of habitats. Preserved_Specimen Baker, R.F. 81 Oahu BISH 1975-04-25
78 Centella asiatica Near mouth of Waipio Stream, Waipio Valley, Big Island, Hawaii Growing on edge of outflow to the stream, from Cyperus papyrus pond area, in sandy wet soil. Preserved_Specimen Crago, L.M. 2005115 Hawaii BISH 2005-03-19
79 Centella asiatica Wailau Valley, n. coast East Molokai Rocky flat above tide level. Preserved_Specimen Fosberg, F.R. 9649 Molokai BISH 1933-07-04
80 Centella asiatica Mokuauia Island, Laie In crevices in coral rock. Preserved_Specimen Fosberg, F.R. 14169 Oahu BISH 1937-07-04
81 Centella asiatica Tantalus, Koolau Mts Wet forest, along trail - disturbed ground. Preserved_Specimen Stemmermann, R.L. 426 Oahu BISH 1973-08-09
82 Centella asiatica Located down Kona-Hilo Hwy 4 mi past Hongo Store towards Hilo; turn right onto Fern Estates Road. Plant was about 200 m down Fern Estates Road, on right side. (plot 55) Scrub forest community. Preserved_Specimen Newell, C.L. 671 Hawaii BISH 1966-08-31
83 Centella asiatica Honolulu. Crest of Koolau Range, near Puu Konahuanui, ca. 3.5 air mi NNE of Univ. of Hawaii campus Native forest; along trail. Preserved_Specimen Young, D.A. 1639 Oahu BISH 1979-01-18
84 Centella asiatica Honolulu: Manoa Valley; Lyon Arboretum-Middle Lawn Makai Collected under eaves of new Greenhouse. Preserved_Specimen Nagata, K.M. 439 Oahu BISH 1968-06-28
85 Centella asiatica Hawaii National Park, Kilauea, Crater Rim Road, near Fern Jungle Trail Locally abundant. Preserved_Specimen Fagerlund, G.O. 1080 Hawaii BISH 1945-06-13
86 Centella asiatica Ditch Trail Kahana Valley, Koolau Mts. Kahana Wet forest, hanging over bank along tract. Preserved_Specimen Fosberg, F.R. 10410 Oahu BISH 1933-11-26
87 Centella asiatica Along Schofield-Waikane Trail, leeward of Koolau Ridge Abundant on moist roadside bank and along trail. Preserved_Specimen Crosby, M.R. 1594 Oahu BISH 1964-06-30
88 Centella asiatica Koolau Range, Manoa; Ridge trail back of Woodlawn Growing on side of tree. Preserved_Specimen Kuykendall, J.R. 66 Oahu BISH 1942-12-31
89 Centella asiatica Lyon Arboretum, Honolulu; Upper Lawn Makai Around base of palms near gate. Very common weed. Preserved_Specimen Baker, R.F. 81 Oahu BISH 1975-04-25
90 Centella asiatica Makawao District, Koolau Forest Reserve, N slope of Haleakala. Small gulch below dilapidated flume, 0.2 mi E of Waikamoi Stream Muddy streambed in wet forest. Preserved_Specimen Lammers, T.G. 5655 Maui BISH 1984-07-13
91 Centella asiatica Lanai City Weed in a lawn. Preserved_Specimen Hobdy, R.W. 2547 Lanai BISH 1986-03-09
92 Centella asiatica So. ridge So. Opaeula Gulch, Koolau Mts. Paalaa Roadside mud. Preserved_Specimen Fosberg, F.R. 8738 Oahu BISH 1932-09-25
93 Centella asiatica Pupukea Forest, Koolauloa Creek bed Aleurites Forest. Preserved_Specimen Grant, M.L. 7154 Oahu BISH 1934-08-01
94 Centella asiatica Koloa District, Lawai Valley, National Tropical Botanical Garden Common in moist, shady areas of valley. Preserved_Specimen Herbst, D.R. 2286 Kauai BISH 1972-01-03
95 Centella asiatica Nuuanu Water Shed, site of ruins of King Kamehameha III's summer home Preserved_Specimen Mill, S.W. 13 Oahu BISH 1983-08-05
96 Centella asiatica Aiea In pineapple field. Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
97 Centella asiatica Waianae Ridge, Maunauna, SE slope In Acacia koa forest. Preserved_Specimen van Royen, P. 10126 Oahu BISH 1967-07-04
98 Centella asiatica Lahaina District, Road to Hailau [probably] on Kaulalewelewe between Honokowai and Mahinahina Valleys Disturbed mesic forest Preserved_Specimen Ray, G. 1062 Maui BISH 1999-02-18
99 Centella asiatica open wet forest Preserved_Specimen Allison, A. Oahu BISH 2021-01-17
100 Centella asiatica Top of pali overlooking leper settlement, Kalaupapa Shade of bush. Preserved_Specimen St.John, H. 12540 Molokai BISH 1932-12-24
101 Centella asiatica East Maui, Hana Rain Forest Project In open bog with Carex svenonis and Erechtites valerianifolia. Preserved_Specimen Harrison, B. 316 Maui BISH 1973-07-09
102 Centella asiatica Kalaupapa; Papapaiki, coast Preserved_Specimen Wysong, M.L. 581 Molokai BISH 2005-01-12
103 Centella asiatica Honolulu, Manoa Valley, Manoa Arboretum Preserved_Specimen Russell, P. 23 Oahu BISH 1932-07-05
104 Centella asiatica Honolulu, Bishop Museum campus Preserved_Specimen Brown, F.B.H. 1323 Oahu BISH 1925-01-03
105 Centella asiatica Nuuanu Valley Preserved_Specimen Forbes, C.N. 1421 Oahu BISH 1909-12-12
106 Centella asiatica West of Waialua, near irrigation ditch, edge of cane field Preserved_Specimen Kerr, M. Oahu BISH 1946-06-30
107 Centella asiatica Manoa Arboretum Preserved_Specimen Caum, E.L. Oahu BISH 1938-07-07
108 Centella asiatica Mokolii Preserved_Specimen Neal, M.C. Oahu BISH 1938-09-24
109 Centella asiatica near the U.S. Forest Service Experimental Tree Planting Site within Halepuaa Forest Reserve in the Puna District Preserved_Specimen Clarke, G. 79.217 Hawaii BISH 1979-03-08
110 Centella asiatica Kaneohe 47-775 Ahilama Rd. (private land holding) Preserved_Specimen Annable, C.R. 3361 Oahu BISH 1997-03-15
111 Centella asiatica ahupuaa of Halepuaa in the Puna District, in the area of teh experimental tree planting site on the Puna Trail OBSERVATION Curtis Daehler Hawaii HAPI 7/19/2007
112 Centella asiatica Oahu Is., koolau range, poamoho trail OBSERVATION Curtis Daehler Oahu HAPI 3/18/2007
113 Centella asiatica Makawaon suojelualue, Kula, HI, US OBSERVATION Curtis Daehler Hawaii HAPI 3/18/2007
114 Centella asiatica Kailua, Ulumawao Ridge Somewhat common on ridges, beneath Osteomeles. Preserved_Specimen Davis, J. 90 Oahu BISH 1979-03-07
115 Centella asiatica Windward side of island, Anapuhi to Wainene, long coastal boulder-strewn beach with vertical cliff at back 10--30 m from water Cliff with Artemisia australis, Chenopodium oahuense, Schiedea globosa, Kalanchoe pinnata, Tetramolopium sylvae, Lythrum maritimum, and Hedyotis littoralis, among others. Preserved_Specimen Wagner, W.L. 5746 Molokai BISH 1987-08-01
116 Centella asiatica Waianae Mts., Puu Manawahua Common ground cover in woods. Preserved_Specimen St.John, H. 9896 Oahu BISH 1929-09-29
117 Centella asiatica Kalepa Ridge Windward side of northern part of ridge. Preserved_Specimen Corn, C. Kauai BISH 1977-07-20
118 Centella asiatica Lanai City, S section Weed in waste area. Preserved_Specimen Nagata, K.M. 3493 Lanai BISH 1986-06-02
119 Centella asiatica East Maui, Hana District, Waihoi Valley Found in marshy depression; seemed to occur in wet disturbed areas with several introduced grasses and sedges. Preserved_Specimen Harrison, B. 25 Maui BISH 1972-09-21
120 Centella asiatica Hilo District, Onomea Valley, Hawaii Tropical Botanical Garden grounds Creeping on ground in deep shade beneath coconut palms, Pandanus trees, and Alexander palms; exposed to salt spray. Preserved_Specimen Staples, G.W. 529 Hawaii BISH 1990-01-08
121 Centella asiatica [Hawaiian Islands; U.S.A.; Molokai; Hawaii; Maui: Molokai: Lanikula Anele was collected at Lanikula under the kukui trees. There were many kukui seedlings, noni seedlings, wiki, and kukae-pua`a among other plants in the beautiful kukui grove. Preserved_Specimen Funk, E.J. 920052 Molokai BISH 1992-05-03
122 Centella asiatica Koolau Mts., Pupukea - along summit trail On grassy summit. Observed several Pritchardia stands. Preserved_Specimen Stone, B.C. 2815 Oahu BISH 1959-05-04
123 Centella asiatica Between Waiahole and Waikane valleys, windward side of Koolau Range On wind-swept ridge. Preserved_Specimen Harris, J.A. 242047 Oahu BISH 1924-08-16
124 Centella asiatica ahupuaa of Halepuaa in the Puna District, in the area of teh experimental tree planting site on the Puna Trail The soil in the area was crushed lava or cinder. The plant was under tall grasses that shielded it from the direct sunlight. Preserved_Specimen Yoshida, L. 79.066 Hawaii BISH 1979-03-08
125 Centella asiatica Waikane-Schofield Trail, Koolau Mts., Waikane Wet trail-side bank. Preserved_Specimen Fosberg, F.R. 8783 Oahu BISH 1932-10-16
126 Centella asiatica Wailua Game Reserve, above Wailua Vegetation on plots stripmined for bauxite ore in 1958-60. Tree plot area. Preserved_Specimen Howard, R.A. 20251 Kauai BISH 1985-10-20
127 Centella asiatica Koloa District. Lower slope of Mt. Kahili off the TV Relay-Alexander Dam Rd., ca. 1.5 mi above the Wahiawa Bog Associated with Hibiscus kahilii, Clermontia clermontioides, Psychotria, Dubautia, Psidium, Metrosideros polymorpha, etc. Preserved_Specimen Gustafson, R. 1056 Kauai BISH 1978-11-25
128 Centella asiatica Waikiki District, Honolulu Vacant lot. Preserved_Specimen Baker, R.J. 228 Oahu BISH 1932-07-20
129 Centella asiatica East Maui, Waihoi Valley, Transect 1 In wet open disturbed areas with Paspalum conjugatum. Preserved_Specimen Harrison, B. 20 Maui BISH 1972-06-15
130 Centella asiatica Kipapa Trail, ~100 m from refuge boundary, along edges of trail wet lowland forest; Metrosideros, Acacia, Dicranopteris Preserved_Specimen Reynolds, L.S. 72 Oahu BISH 2006-05-31
131 Centella asiatica Lyon Arboretum, Honolulu; Upper Lawn Makai Around base of palms near gate. Very common weed. Wide range of habitats. Preserved_Specimen Baker, R.F. 81 Oahu BISH 1975-04-25
132 Centella asiatica Near mouth of Waipio Stream, Waipio Valley, Big Island, Hawaii Growing on edge of outflow to the stream, from Cyperus papyrus pond area, in sandy wet soil. Preserved_Specimen Crago, L.M. 2005115 Hawaii BISH 2005-03-19
133 Centella asiatica Wailau Valley, n. coast East Molokai Rocky flat above tide level. Preserved_Specimen Fosberg, F.R. 9649 Molokai BISH 1933-07-04
134 Centella asiatica Mokuauia Island, Laie In crevices in coral rock. Preserved_Specimen Fosberg, F.R. 14169 Oahu BISH 1937-07-04
135 Centella asiatica Tantalus, Koolau Mts Wet forest, along trail - disturbed ground. Preserved_Specimen Stemmermann, R.L. 426 Oahu BISH 1973-08-09
136 Centella asiatica Located down Kona-Hilo Hwy 4 mi past Hongo Store towards Hilo; turn right onto Fern Estates Road. Plant was about 200 m down Fern Estates Road, on right side. (plot 55) Scrub forest community. Preserved_Specimen Newell, C.L. 671 Hawaii BISH 1966-08-31
137 Centella asiatica Honolulu. Crest of Koolau Range, near Puu Konahuanui, ca. 3.5 air mi NNE of Univ. of Hawaii campus Native forest; along trail. Preserved_Specimen Young, D.A. 1639 Oahu BISH 1979-01-18
138 Centella asiatica Honolulu: Manoa Valley; Lyon Arboretum-Middle Lawn Makai Collected under eaves of new Greenhouse. Preserved_Specimen Nagata, K.M. 439 Oahu BISH 1968-06-28
139 Centella asiatica Hawaii National Park, Kilauea, Crater Rim Road, near Fern Jungle Trail Locally abundant. Preserved_Specimen Fagerlund, G.O. 1080 Hawaii BISH 1945-06-13
140 Centella asiatica Ditch Trail Kahana Valley, Koolau Mts. Kahana Wet forest, hanging over bank along tract. Preserved_Specimen Fosberg, F.R. 10410 Oahu BISH 1933-11-26
141 Centella asiatica Along Schofield-Waikane Trail, leeward of Koolau Ridge Abundant on moist roadside bank and along trail. Preserved_Specimen Crosby, M.R. 1594 Oahu BISH 1964-06-30
142 Centella asiatica Koolau Range, Manoa; Ridge trail back of Woodlawn Growing on side of tree. Preserved_Specimen Kuykendall, J.R. 66 Oahu BISH 1942-12-31
143 Centella asiatica Lyon Arboretum, Honolulu; Upper Lawn Makai Around base of palms near gate. Very common weed. Preserved_Specimen Baker, R.F. 81 Oahu BISH 1975-04-25
144 Centella asiatica Makawao District, Koolau Forest Reserve, N slope of Haleakala. Small gulch below dilapidated flume, 0.2 mi E of Waikamoi Stream Muddy streambed in wet forest. Preserved_Specimen Lammers, T.G. 5655 Maui BISH 1984-07-13
145 Centella asiatica Lanai City Weed in a lawn. Preserved_Specimen Hobdy, R.W. 2547 Lanai BISH 1986-03-09
146 Centella asiatica So. ridge So. Opaeula Gulch, Koolau Mts. Paalaa Roadside mud. Preserved_Specimen Fosberg, F.R. 8738 Oahu BISH 1932-09-25
147 Centella asiatica Pupukea Forest, Koolauloa Creek bed Aleurites Forest. Preserved_Specimen Grant, M.L. 7154 Oahu BISH 1934-08-01
148 Centella asiatica Koloa District, Lawai Valley, National Tropical Botanical Garden Common in moist, shady areas of valley. Preserved_Specimen Herbst, D.R. 2286 Kauai BISH 1972-01-03
149 Centella asiatica Nuuanu Water Shed, site of ruins of King Kamehameha III's summer home Preserved_Specimen Mill, S.W. 13 Oahu BISH 1983-08-05
150 Centella asiatica Aiea In pineapple field. Preserved_Specimen Christophersen, E. Oahu BISH 1924-12-07
151 Centella asiatica Waianae Ridge, Maunauna, SE slope In Acacia koa forest. Preserved_Specimen van Royen, P. 10126 Oahu BISH 1967-07-04
152 Centella asiatica Lahaina District, Road to Hailau [probably] on Kaulalewelewe between Honokowai and Mahinahina Valleys Disturbed mesic forest Preserved_Specimen Ray, G. 1062 Maui BISH 1999-02-18
153 Centella asiatica open wet forest Preserved_Specimen Allison, A. Oahu BISH 2021-01-17
154 Centella asiatica Top of pali overlooking leper settlement, Kalaupapa Shade of bush. Preserved_Specimen St.John, H. 12540 Molokai BISH 1932-12-24
155 Centella asiatica East Maui, Hana Rain Forest Project In open bog with Carex svenonis and Erechtites valerianifolia. Preserved_Specimen Harrison, B. 316 Maui BISH 1973-07-09
156 Centella asiatica Kalaupapa; Papapaiki, coast Preserved_Specimen Wysong, M.L. 581 Molokai BISH 2005-01-12
157 Centella asiatica Honolulu, Manoa Valley, Manoa Arboretum Preserved_Specimen Russell, P. 23 Oahu BISH 1932-07-05
158 Centella asiatica Honolulu, Bishop Museum campus Preserved_Specimen Brown, F.B.H. 1323 Oahu BISH 1925-01-03
159 Centella asiatica Nuuanu Valley Preserved_Specimen Forbes, C.N. 1421 Oahu BISH 1909-12-12
160 Centella asiatica West of Waialua, near irrigation ditch, edge of cane field Preserved_Specimen Kerr, M. Oahu BISH 1946-06-30
161 Centella asiatica Manoa Arboretum Preserved_Specimen Caum, E.L. Oahu BISH 1938-07-07
162 Centella asiatica Mokolii Preserved_Specimen Neal, M.C. Oahu BISH 1938-09-24
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